Hemiphyllodactylus ziegleri sp. nov. (Squamata, Gekkonidae), a new karst-dwelling gecko species from Son La Province, Vietnam Author Pham, Anh Van 0000-0002-6023-3418 Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam Author Nguyen, Truong Quang 0000-0002-6601-0880 Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 10072, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 11416, Vietnam Author Pham, Cuong The 0000-0001-5158-4526 Institute of Biology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 10072, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi 11416, Vietnam Author Ngo, Hanh Thi 0000-0002-5283-6243 Central Institute for Natural Resources and Environmental Studies, Vietnam National University, Hanoi, 19 Le Thanh Tong, Hanoi 11021, Vietnam Author Le, Minh Duc 0000-0002-2953-2815 Faculty of Environmental Sciences, University of Science, Vietnam National University, Hanoi, 334 Nguyen Trai Road, Hanoi, Vietnam & Department of Herpetology, American Museum of Natural History, Central Park West at 79 th Street, New York 10024 USA text ZooKeys 2026 2026-02-04 1268 75 94 journal article 314137 10.3897/zookeys.1268.174678 cc2d4030-e507-4e5b-9770-e7760d510787 EFE111E7-A52A-4FF4-BB38-0526245DA4D2

Table 2. Morphological characters of Hemiphyllodactylus ziegleri sp. nov. from Son La Province, Vietnam (measurements in mm, * = regenerated or broken tail, min = minimum, max = maximum, other abbreviations defined in the text).

Characters	HUS.2025.06	HUS.2025.08	IB R.6431	HUS.2025.07	IB R.6433	Min-max	HUS.2025.09	HUS.2025.10	IB R.6432	HUS.2025.11	Min-max
	Holotype	Paratype	Paratype	Paratype	Paratype		Paratype	Paratype	Paratype	Paratype
Sex	M	M	M	M	M		F	F	F	F
SVL	41.9	41.8	41.3	38.7	40	38.7–41.9	49.3	47.2	47	47.5	47–49.3
TaL	12.6 *	18.6 *	32.64	26.5 *	26.9 *		30.18 *	28.7 *	31.21	36.0	31.21–36
TrunkL	20.08	21.8	20.68	19.5	20.3	19.5–21.8	23.8	23	23.2	24.05	23–24.05
EyeD	2.68	2.62	2.56	2.55	2.6	2.55–2.68	3.23	3.11	3.09	2.9	2.9–3.23
HeadL	9.62	9.67	9.69	9.32	9.54	9.32–9.69	11.5	11.35	11.26	10.6	10.6–11.5
HeadW	7.24	7.08	7.22	6.94	7.02	6.94–7.24	9.1	9	8.9	7.96	7.96–9.1
Nare-eye length (NarEye)	3.24	3.2	3.22	3.2	3.29	3.2–3.29	4.09	3.94	3.92	3.5	3.5–4.09
Snout-eye length (SnEye)	4.51	4.59	4.58	4.3	4.45	4.3–4.59	5.45	5.44	5.15	5.06	5.06–5.45
Internarial distance (SnW)	1.59	1.75	1.77	1.48	1.6	1.48–1.77	2.06	2.12	1.89	1.96	1.89–2.12
Ear opning diameter (EarD)	0.91	0.96	0.92	0.9	0.97	0.9–0.97	1.11	1.12	1.08	1.06	1.06–1.12
EarD / EyeD	0.34	0.37	0.36	0.35	0.37	0.34–0.37	0.34	0.36	0.35	0.37	0.34–0.37
Trunk / SVL	0.48	0.52	0.50	0.50	0.51	0.48–0.52	0.48	0.49	0.49	0.51	0.48–0.51
HeadL / SVL	0.23	0.23	0.23	0.24	0.24	0.23–0.24	0.23	0.24	0.24	0.22	0.22–0.24
HeadW / SVL	0.17	0.17	0.17	0.18	0.18	0.17–0.18	0.18	0.19	0.19	0.17	0.17–0.19
HeadW / HeadL	0.75	0.73	0.75	0.74	0.74	0.73–0.75	0.79	0.79	0.79	0.75	0.75–0.79
SnEye / HeadL	0.47	0.47	0.47	0.46	0.47	0.46–0.47	0.47	0.48	0.46	0.48	0.46–0.48
NarEye / HeadL	0.34	0.33	0.33	0.34	0.34	0.33–0.34	0.36	0.35	0.35	0.33	0.33–0.36
EyeD / Head L	0.28	0.27	0.26	0.27	0.27	0.26–0.28	0.28	0.27	0.27	0.27	0.27–0.28
SnW / Hea dL	0.17	0.18	0.18	0.16	0.17	0.16–0.18	0.18	0.19	0.17	0.18	0.17–0.19
EyeD / NarEye	0.83	0.82	0.80	0.80	0.79	0.79–0.83	0.79	0.79	0.79	0.83	0.79–0.83
SnW / HeadW	0.22	0.25	0.25	0.21	0.23	0.21–0.25	0.23	0.24	0.21	0.25	0.21–0.25
Dorsal scale row s	22–23	25–27	23–24	25–26	22–23	22–27	21–22	22–24	22	23–24	21–24
Ventral scale row s	12	13	14	13	14	12–14	16	13	13	16	13–16
Cloacal spurs (CloacS)	1	1	1	1	1	1–1	1	1	1	1	1–1
Circumnasal (CircN)	3	3	3	3	3	3–3	3	3	3	3	3–3
Scales between supranasals (SnS)	3	3	3	3	3	3–3	3	2	3	1	1–3
Supralabials (Sublab)	10 / 10	10 / 10	10 / 10	11 / 10	10 / 10	10–11	12 / 12	12 / 11	11 / 12	11 / 11	11–12
Infralabials (Inflab)	9 / 9	9 / 9	9 / 9	9 / 9	9 / 9	9	10 / 10	9 / 9	9 / 10	10 / 10	9–10
Chin scale (Chin)	4	7	7	7	4	4–7	7	7	7	7	7
Lamella formula
forelimbs (pairs only)	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4	3-4 - 4 - 4
hindlimbs (pairs only)	4-4 - 4 - 4	4-4 - 4 - 4	4-5 - 5 - 5	4-4 - 5 - 4	4-5 - 5 - 5 - 5	4-4 / 5 - 4 / 5 - 4 / 5	4-5 - 5 - 5	4-5 - 5 - 5	4-5 - 5 - 5	4-4 - 5 - 4	4-4 / 5 - 4 / 5 - 4 / 5
First digit
forelimbs	5	4	3	4	5	3–5	5	5	5	4	4–5
hindlimbs	6	5	5	5	5	5–6	4	5	5	4	4–5
Precloacal pores	21	21	21	21	23	21–23	0	0	0	0	0