| Eye diameter (% HL) | 6.8–9.1 | 6.1–9.2 | 6.2–7.1 | 5.2–6.7 | 6.0–7.9 | 5.1–7.9 | 4.1–6.9 |
| Interorbital distance (% HL) | 24.1–28.8 | 22.8–29.6 | 20.4–23.8 | 23.4–26.0 | 22.4–25.9 | 25.4–26.8 | 27.1–30.0 |
| Head length behind the eyes (% HL) | 35.6–40.3 | 35.4–39.4 | 35.8–42.7 | 40.4–45.5 | 34.7–40.6 | —— | —— |
| Maxillary barbel length (% HL) | 67.1–84.8 | 65.9–82.1 | 73.5–79.6 | 73.8–84.0 | 77.1–90.3 | 90.5–97.7 | 91.1–96.2 |
| Head depth (% HL) | 38.9–53.1 | 42.0–48.4 | 37.2–41.1 | 36.4–47.6 | 34.8–52.1 | —— | —— |
| Head width (% SL) | 18.5–24.5 | 18.1–21.0 | 18.0–21.2 | 19.9–22.8 | 19.6–21.8 | 21.3–23.9 | 20.6–23.5 |
| Caudal-peduncle length (% SL) | 21.5–29.2 | 20.6–26.7 | 21.2–26.1 | 18.7–24.3 | 22.3–29.2 | 21.0–25.0 | 15.7–19.6 |
| Dorsal-fin insertion to adipose-fin origin distance (% SL) | 14.1–23.2 | 15.5–25.5 | 12.1–25.1 | 15.3–25.7 | 12.8–20.1 | 26.5–33.5 | 25.2–28.4 |
| Pelvic-fin origin to anal-fin origin distance (% SL) | 24.4–32.4 | 27.8–33.1 | 27.2–31.6 | 29.2–32.5 | 24.1–28.6 | —— | —— |
| Pelvic insertion to adipose origin distance (% SL) | 15.4–19.5 | 17.3–22.3 | 13.8–21.3 | 16.3–21.3 | 10.5–17.0 | —— | —— |
| Post-adipose length (% SL) | 8.2–12.7 | 8.1–11.2 | 7.6–10.4 | 6.7–10.2 | 7.3–13.7 | 7.2–9.8 | 4.6–7.6 |
| Caudal-peduncle depth (% caudal-peduncle of length) | 27.1–40.3 | 25.9–40.6 | 32.7–41.5 | 21.8–41.3 | 13.0–24.6 | 21.9–28.1 | 37.8–49.2 |
| Pectoral-fin depth (% pectoral-fin length) | 53.0–70.5 | 61.0–78.2 | 38.4–51.4 | 42.2–56.3 | 51.0–72.7 | —— | —— |
| Pelvic-fin width (% pelvic-fin length) | 43.0–68.4 | 56.6–76.6 | 35.6–49.0 | 42.0–60.0 | 42.7–74.4 | —— | —— |
| Dorsal-fin ray | i, 5 1 / 2 | i, 5 1 / 2 | i, 4 1 / 2 | i, 4 1 / 2 | i, 5 1 / 2 | i, 5 1 / 2 | i, 5 1 / 2 |
| Pectoral-fin ray | i, 12–13 | i, 12–14 | i, 13–14 | i, 13–14 | i, 13–14 | i, 14–15 | i, 13–14 |
| Anal-fin ray | i, 4 1 / 2 | i, 4 1 / 2 | i, 3 1 / 2 | i, 3 1 / 2 | i, 3 1 / 2 | i, 4 1 / 2 | i, 4 1 / 2 |
| Caudal-fin ray | i, 7 + 8, i | i, 7 + 8, i | i, 7 + 8, i | i, 6 + 7, i | i, 6 + 7, i | i, 7 + 8, i | i, 7 + 8, i |
| Vertebra number | 13 + 25 = 38 (n = 1); 13 + 24 = 37 (n = 1); 12 + 24 = 36 (n = 2) | 13 + 23 = 36 (n = 2); 13 + 24 = 37 (n = 2) | 13 + 24 = 37 (n = 1); 13 + 23 = 36 (n = 1); 13 + 25 = 38 (n = 2) | 12 + 21 = 33 (n = 1); 12 + 20 = 32 (n = 1); 11 + 21 = 32 (n = 2) | 12 + 22 = 34 (n = 2); 13 + 21 = 34 (n = 1); 12 + 21 = 33 (n = 1) | 12 + 21 = 33 (n = 1) | 19 + 15 = 34 (n = 2) |
| Lower corner of gill opening | Opposite base of the second and third pectoral-fin elements | Opposite base of the fourth and fifth pectoral-fin elements | Opposite base of the fourth and fifth pectoral-fin elements | Opposite base of the second and third pectoral-fin elements | Opposite base of the second and third pectoral-fin elements | Opposite base of the fourth and fifth pectoral-fin elements | Opposite base of the second and third pectoral-fin elements |
| Maxillary barbel tip | Not reaching or extending beyond lower gill-opening corner | Not reaching lower gill-opening corner | Extending beyond lower gill-opening corner | Not reaching or reaching lower gill-opening corner | Reaching or extending beyond lower gill-opening corner | Not reaching lower gill-opening corner | Reaching lower gill-opening corner |
| Nasal barbel tip | Not reaching anterior orbital | Not reaching anterior orbital | Extending beyond anterior orbital | Extending beyond anterior orbital and not reaching posterior orbital | Not reaching or reaching anterior orbital | Not reaching anterior orbital | Extending beyond posterior orbital |
| Pelvic-fin tip | Reaching or extending beyond anus | Not reaching, reaching, or extending beyond anus | Not reaching anus | Reaching or extending beyond anus | Reaching or extending beyond anus | Not reaching or reaching anus (♀); extending beyond anus (♂) | Not reaching anus (♀); reaching anus (♂) |
| Ventral abdominal surface | convex | flattened | flattened | convex | flattened | flattened | – |