Two new species of semiaquatic Anolis (Squamata: Dactyloidae) from Costa Rica Author Chaves, Gerardo 0000-0002-4301-6569 Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & cachi 13 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4301 - 6569 cachi13@gmail.com Author Ryan, Mason J. Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA & Arizona Game and Fish Department, 5000 W Carefree Hwy, Phoenix, Arizona 85086 Author Bolaños, Federico 0000-0002-7935-6418 Centro de Investigación en Biodiversidad y Ecología Tropical (Museo de Zoología) de la Universidad de Costa Rica & Escuela de Biología, Universidad de Costa Rica, San Pedro, Costa Rica & federico. bolanos @ ucr. ac. cr; https: // orcid. org / 0000 - 0002 - 7935 - 6418 federico.bolanos@ucr.ac.cr Author Márquez, Cruz 0009-0003-2220-7379 Direccion, Conservacion, Restauracion y Desarrollo Sustentable de Ecosistemas Insulares; Parque Nacional Galapagos, Ecuador cruzkallw @ gmail. com; https: // orcid. org / 0009 - 0003 - 2220 - 7379 cruzkallw@gmail.com Author Köhler, Gunther 0000-0002-2563-5331 Senckenberg Forschungsinstitut und Naturmuseum, Senckenberganlage 25, 60325 Frankfurt a. M., Germany gkoehler @ senckenberg. de; https: // orcid. org / 0000 - 0002 - 2563 - 5331 gkoehler@senckenberg.de Author Poe, Steven Department of Biology, University of New Mexico, Albuquerque, NM, 87131, USA text Zootaxa 2023 2023-07-25 5319 2 249 262 http://dx.doi.org/10.11646/zootaxa.5319.2.6 journal article 57778 10.11646/zootaxa.5319.2.6 49b790fb-5325-46da-aa80-d36d2e1fcc1a 1175-5326 8182539 065A967F-2832-4BDF-890A-2EA852CACD74

TABLE 1. Selected measurements and scale characters of Anolis aquaticus, A. riparius, and A. robinsoni. Mean value

VARIABLES	Anolis aquaticus	Anolis riparius	Anolis robinsoni	MANOVA
and one standard deviation are followed by range in parentheses. MANOVA results are for selected variables and all with two degrees of freedom.
Number of individuals	35	8	29
Dewlap coloration	Yellow and red	Yellow and red	Dark brown
SVL (mm)	57.0 ± 8.5 (30.6–69.5)	60.8 ± 7.2 (30.6–73.3)	58.9 ± 9.9 (23.1–73.8)
Extent of dewlap past axilla in males (mm)	6.4 ± 2.5 (0.1–11.3)	8.2 ± 3.0 (6.1–10.3)	8.8 ± 2.7 (3.5–12.2)
HL	15.6 ± 1.9 (11.4–19.5)	15.4 ± 2.6 (9.6–20.1)	15.0 ± 2.6 (8.2–18.8)	F = 0.35, P= 0.35
HW	9.2 ± 1.1 (6.4–10.7)	9.5 ± 1.4 (5.9–12.4)	9.3 ± 2.2 (4.7–18.2)	F = 0.07, P= 0.92
Small caudal scales before the first larger scales	11 ± 1 (9–14)	18 ± 2 (16–21)	18 ± 1 (15–20)	F = 52.05, P<0.01
Scales between supraocular semicircles	4 ± 1 (2–7)	4 ± 0 (4–5)	6 ± 1 (4–7)	F = 17.08, P<0.01
Scales between interparietal and supraoculars semicircle	6 ± 1 (5–10)	9 ± 1 (7–11)	7 ± 1 (5–11)	F = 30.04, P<0.01
Supralabials	9 ± 1 (7–11)	11 ± 1 (10–12)	9 ± 1 (7–11)	F = 12.43, P<0.01
Infralabials	9 ± 1 (8–10)	11 ± 1 (10–12)	9 ± 1 (8–11)	F = 7.17, P<0.01
Lamellae under fourth toe	14 ± 1 (11–16)	15 ± 1 (14–17)	16 ± 1 (15–17)	F = 12.44, P<0.01
Loreal scales	9 ± 1 (7–11)	9 ± 1 (8–10)	9 ± 1 (8–11)
Scales between nasal and rostral	1 ± 0 (0–2)	1 ± 1 (1–2)	2 ± 0 (1–3)
SO/LA	2 ± 0 (1–2)	1 ± 1 (1–2)	1 ± 0 (1–2)
Postrostral scales	9 ± 1 (7–11)	10 ± 1 (8–11)	9 ± 1 (7–10)	F = 0.59, P= 0.55