French Mediterranean islands as a refuge of relic earthworm species: Cataladrilus porquerollensis sp. nov. and Scherotheca portcrosana sp. nov. (Crassiclitellata, Lumbricidae) Author Marchán, Daniel F. 3B3731B6-B5FB-409A-A7A3-99FD0F96D688&38538B17-F127-4438-9DE2-F9D6C597D044 Grupo de Ecoloxía Animal (GEA), Universidade de Vigo, E- 36310 Vigo, Spain. CEFE, Univ Montpellier, CNRS, EPHE, IRD, Univ Paul Valéry Montpellier 3, Montpellier, France. Department of Biodiversity, Ecology and Evolution, Faculty of Biology, Universidad Complutense de Madrid, Madrid, Spain. UMR Eco & Sols, INRAE-IRD-CIRAD-SupAgro Montpellier, 2 Place Viala, 34060 Montpellier Cedex 2, France. Taxonomia Biodiversity Fund, 7 rue Beccaria, 72012, Paris, France. dadico@ucm.es Author Decaëns, Thibaud B61F61B2-3012-4526-8FF9-DC94D372AF77 CEFE, Univ Montpellier, CNRS, EPHE, IRD, Univ Paul Valéry Montpellier 3, Montpellier, France thibaud.DECAENS@cefe.cnrs.fr Author Díaz Cosín, Darío J. 38538B17-F127-4438-9DE2-F9D6C597D044 Department of Biodiversity, Ecology and Evolution, Faculty of Biology, Universidad Complutense de Madrid, Madrid, Spain dadico@ucm.es Author Hedde, Mickaël F4A219F7-7E75-4333-8293-3004B3CD62C5 UMR Eco & Sols, INRAE – IRD – CIRAD – SupAgro Montpellier, 2 Place Viala, 34060 Montpellier Cedex 2, France mickael.hedde@inra.fr Author Lapied, Emmanuel B1FB8658-DFC3-481C-A0BE-B8488A018611 Taxonomia Biodiversity Fund, 7 rue Beccaria, 72012, Paris, France elapied@yahoo.com Author Domínguez, Jorge 167575D5-D2CC-4B37-8B1D-0233E6B154E5 Grupo de Ecoloxía Animal (GEA), Universidade de Vigo, E- 36310 Vigo, Spain. CEFE, Univ Montpellier, CNRS, EPHE, IRD, Univ Paul Valéry Montpellier 3, Montpellier, France. Department of Biodiversity, Ecology and Evolution, Faculty of Biology, Universidad Complutense de Madrid, Madrid, Spain. UMR Eco & Sols, INRAE-IRD-CIRAD-SupAgro Montpellier, 2 Place Viala, 34060 Montpellier Cedex 2, France. Taxonomia Biodiversity Fund, 7 rue Beccaria, 72012, Paris, France. jdguez@uvigo.es text European Journal of Taxonomy 2020 2020-08-13 701 1 22 journal article 21296 10.5852/ejt.2020.701 73792db7-0c46-4ef2-9d83-ecd3e3f42935 3988368 D9291955-F619-46EA-90E1-DA756D1B7C55

Table 3. Morphological characters of Cataladrilus porquerollensis Marchán & Decäens sp. nov. and morphologically similar relatives within the genera Cataladrilus Qiu & Bouché, 1998 and Prosellodrilus Bouché, 1972. Length expressed in mm, weight expressed in mg.

Characters	Cataladrilus porquerollensis sp. nov.	Cataladrilus (Latisinella) mrsici	Cataladrilus albus	Prosellodrilus dactylothecus
Length	45–51	35	120–145	92
N. segments	196	168	229–297	207
Weight	140	102	1526	409
Pigmentation	Absent	Absent	Absent	Absent
Prostomium	Epilobous	Epilobous	Epilobous	Epilobous
Setae	Closely paired	Separate	Closely paired	Closely paired
Setal arrangement	9:1:3:1:7	5:5:5:4:8	15:1:5:1:15	20:1:10:1:30
Papillae	11, 13	Absent	28–34	13, 14, 15
First dorsal pore	10/11–11/12	6/7	6/7	5/6
Nephridial pores	Aligned, b	Aligned, b	Aligned, b	Aligned, b
Male pores	½ 15, b	½ 15, b	½ 15, b	⅔ 15
Female pores	½ 14, inconspicuous	½ 14, b	½ 14, b	½ 14, b
Spermathecal pores	9/10, 10/11 simple in c	9/10, 10/11 simple in c	9/10, 10/11 simple in d	13/14, 14/15 in c
Clitelum	(19)20–32, 33(34)	22–32	22–34	19–32
Tubercula pubertatis	(28)29–31	29–31	(½ 29)30–34	22–25
Oesophageal hearts	6–11	6–11	6–11	6–11
Calciferous glands	11–13	11–14, dilated in 12, 13	11–14, diverticles in 11	11–14, diverticles in 11
Crop	15–16	15–16	15–16	15–16
Gizzard	17–18	17–18	17–18	17–18
Typhlosole	Bifid	Simple	Bifid	Bifid
Nephridial bladders	U-shaped, reclinate	U-shaped, reclinate	U-shaped, reclinate	U-shaped, reclinate, proximal ampulla
Seminal vesicles	11, 12	9, 11, 12	11, 12	11, 12
Spermathecae	Simple, globular in 10, 11	Simple, pyriform in 10, 11	Simple, small globular in 10, 11	Simple, pedunculate in 14, 15