Reassessment of species delimitation using nuclear markers in three lentic-breeding salamanders from the Chugoku District of Japan (Amphibia: Caudata: Hynobiidae) Author Tomimori, Yusuke 0009-0001-1122-4611 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu-cho, Sakyo-ku, Kyoto 606 - 8501, JAPAN Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN Oshigome 4 - 42 - 1, Kure, Hiroshima 737 - 0915, JAPAN; bigriver @ kb 3. so-net. ne. jp; https: // orcid. org / 0009 - 0005 - 3074 - 833 X Kuta 435 - 19 - 103, Izuhara-Machi, Tsushima, Nagasaki 817 - 0032, JAPAN & katoi 1104 @ gmail. com; https: // orcid. org / 0009 - 0001 - 1122 - 4611 katoi1104@gmail.com Author Matsui, Masafumi 0000-0003-2032-2528 Graduate School of Human and Environmental Studies, Kyoto University, Yoshida Nihonmatsu-cho, Sakyo-ku, Kyoto 606 - 8501, JAPAN Graduate School of Global Environmental Studies, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto 606 - 8501, JAPAN Oshigome 4 - 42 - 1, Kure, Hiroshima 737 - 0915, JAPAN; bigriver @ kb 3. so-net. ne. jp; https: // orcid. org / 0009 - 0005 - 3074 - 833 X Kuta 435 - 19 - 103, Izuhara-Machi, Tsushima, Nagasaki 817 - 0032, JAPAN & fumi @ zoo. zool. kyoto-u. ac. jp; https: // orcid. org / 0000 - 0003 - 2032 - 2528 fumi@zoo.zool.kyoto-u.ac.jp Author Okawa, Hiroshi 0009-0005-3074-833X Author Nishikawa, Kanto 0000-0002-6274-4959 nishikawa. kanto. 8 v @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0002 - 6274 - 4959 nishikawa.kanto.8v@kyoto-u.ac.jp Author Tanabe, Shingo 0009-0002-4764-1852 hynobiustsurugi @ gmail. com; https: // orcid. org / 0009 - 0002 - 4764 - 1852 hynobiustsurugi@gmail.com Author Kamasaka, Ryo text Zootaxa 2023 2023-05-19 5293 1 145 160 http://dx.doi.org/10.11646/zootaxa.5293.1.6 journal article 53363 10.11646/zootaxa.5293.1.6 61b8d37f-c978-4eb6-bed7-f3872f5f6807 1175-5326 7959833 64BEF4FF-6A18-4D96-A382-3218D9F057D4

TABLE 2. Means±SD of SVL and medians of ratios of metric characters (R = %SVL), CG (L), LO in male.

	H. sumidai (n=19)	H. geiyoensis (n=16)	H. akiensis (n=27)
SVL	48.0±4.6 (36.2–56.9)	60.4±3.7 (53.3–67.3)	48.8±4.4 (40.8–57.5)
RHL	23.7 (22–26.7)	22.3 (20.7–24.8)	23.8 (21.5–25.2)
RHW	17.1 (16.6–18.2)	17.1 (15.8–18.4)	17.4 (15.6–19.2)
RMXHW	18.6 (16.7–20.3)	17.9 (16.0–20.6)	18.3 (16.6–22.4)
RLJL	14.6 (13.1–16.4)	13.9 (12.6–15.1)	14.3 (11.9–17.0)
RSL	7.4 (6.1–9.3)	7.5 (6.8–8.3)	7.8 (6.7–8.8)
RIND	5.7 (5.1–6.9)	6.3 (5.6–7.1)	6.3 (5.0–8.0)
RIOD	6.0 (5.1–7.1)	6.6 (5.4–7.6)	6.4 (5.2–7.9)
RUEW	3.0 (2.6–3.7)	2.6 (2.2–3.1)	3.0 (2.6–3.6)
RUEL	5.9 (5.2–6.6)	5.5 (4.9–5.8)	5.8 (5.0–7.0)
RAGD	50.3 (48.6–55.9)	51.0 (46.4–53.9)	49.9 (43.2–54.8)
RTRL	77.6 (72–79.2)	78.9 (76.2–80.9)	76.8 (75.0–79.8)
RTAL	73.6 (63.8–81.8)	78.7 (73.9–87.5)	76.2 (66.0–84.0)
RBTAW	10.8 (9.3–12.3)	11.1 (9.6–12.1)	11.2 (9.0–13.1)
RMTAW	6.2 (4–7.5)	6.9 (5.4–8.1)	6.6 (4.5–8.5)
RBTAH	11.3 (9.1–14.4)	10.6 (9.3–12.8)	11.3 (8.9–14.1)
RMTAH	11.9 (9.2–14.6)	11.1 (9.5–14.1)	11.4 (9.4–14.3)
RFLL	23.6 (20.2–27.7)	24.7 (21.4–26.1)	25.3 (22.3–28.7)
RHLL	29.5 (25.4–32.5)	30.6 (26.9–32.9)	30.8 (27.8–36.1)
R2FL	5.1 (4–6.5)	5.3 (4.2–6.1)	5.2 (3.9–6.4)
R3FL	4.3 (3.4–5.2)	4.8 (3.5–5.2)	4.3 (2.7–5.4)
R3TL	8.5 (4.8–9.9)	8.6 (7.8–9.8)	8.5 (6.5–10.2)
R5TL	2.0 (0–3.5)	2.7 (1.6–4.1)	1.8 (0–4.0)
RVTW	6.0 (4.6–7.1)	5.4 (4.9–6.0)	5.7 (4.2–7.5)
RVTL	4.9 (4.3–6.3)	4.6 (3.8–5.5)	4.6 (3.8–5.9)
CGN	12 (12–13)	12 (12–13)	12 (11–13)
LON	-1 (-2–0)	-1.3 (-2–0)	-0.5 (-2–0.5)