Two new species of the loach genus Homatula Nichols, 1925 (Pisces, Nemacheilidae) from the upper Chang-Jiang Basin in China Author Cao, Liang 0000-0003-3208-1370 Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, Hubei Province, China Author Liu, Yi Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, Hubei Province, China & University of Chinese Academy of Sciences, Beijing 100049, China Author Zeng, Zhixuan 0000-0001-9740-1342 The Department of Endocrinology, Hubei Clinical Medical Research Center for Endocrinology and Metabolic Diseases, Branch of National Clinical Research Center for Metabolic Diseases, Tongji Hospital, Huazhong University of Science and Technology, Wuhan 430030, Hubei Province, China Author Yi, Wenjing Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, Hubei Province, China Author Zhang, E 0000-0002-6971-7160 Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, Hubei Province, China text Zoosystematics and Evolution 2025 2025-03-26 101 2 697 710 journal article 308812 10.3897/zse.101.146027 17a8eb26-f9d7-4b55-a517-86009c547add B49514A4-5753-4411-8156-4C43D6788B8F

Table 3. Genetic distances of Cyt b amongst species of Homatula.

Species		1	2	3	4	5	6	7	8	9	10	11	12	13	14	15	16	17	18	19	20	21	22	23	24
H. shexiang sp. nov.	1
H. xiangzhi sp. nov.	2	3.6
H. tigris	3	6.7	7.9
H. variegata	4	7.8	7.8	8.5
H. berezowskii	5	7.7	8.0	8.0	3.2
H. guanhensis	6	7.6	7.7	8.4	1.4	3.0
H. laxiclathra	7	6.8	7.3	8.0	4.8	4.6	4.6
H. oxygnathra	8	7.8	7.7	8.1	5.3	4.9	5.1	2.1
H. longidorsalis	9	6.7	7.2	7.4	5.7	5.5	5.7	4.7	5.1
H. nanpanjiangensis	10	7.4	7.1	6.7	5.1	5.1	5.2	4.4	4.8	4.5
H. wujiangensis	11	8.5	8.9	10.0	9.3	8.9	9.3	8.9	9.3	8.7	8.2
H. potanini	12	8.6	8.4	10.2	8.8	8.8	8.7	8.8	9.2	8.6	8.1	1.1
H. anguillioides	13	8.2	8.7	8.2	9.7	9.4	9.5	9.1	9.3	8.4	7.8	9.7	9.5
H. acuticephala	14	8.4	8.6	8.3	9.6	9.3	9.4	9.0	9.2	8.3	7.7	9.5	9.4	0.1
H. anteridorsalis	15	10.3	10.1	10.8	11.0	11.1	10.6	10.5	10.4	9.9	9.1	11.2	10.9	4.1	4.0
H. cryptoclathrata	16	9.8	9.8	10.4	10.6	10.5	10.1	9.7	9.7	9.5	8.8	10.6	10.5	3.8	3.7	1.8
H. disparizona	17	8.1	8.6	9.0	9.8	9.4	10.0	8.4	8.8	8.2	7.5	9.7	9.8	8.5	8.6	10.0	9.3
H. geminusclathrata	18	12.5	11.5	12.9	14.8	14.2	14.5	14.6	14.4	13.0	13.0	13.2	13.0	11.9	12.0	13.0	12.9	13.2
H. longibarbata	19	8.8	8.3	9.6	10.5	9.9	10.1	9.5	9.2	9.2	8.3	9.3	9.1	4.2	4.1	4.2	4.3	9.8	11.6
H. microcephala	20	10.2	10.3	10.0	11.0	10.8	11.1	11.0	10.9	9.8	9.6	10.3	10.3	4.5	4.5	4.2	4.1	9.5	11.6	4.9
H. nigra	21	9.8	10.0	10.1	10.4	10.4	10.3	9.8	9.6	9.2	8.9	10.5	10.2	4.2	4.2	2.4	1.5	8.8	11.6	4.0	2.7
H. pycnolepis	22	9.0	9.8	8.8	10.5	10.1	10.2	9.5	9.9	8.9	8.4	9.8	9.8	3.3	3.4	4.0	3.9	8.6	11.4	4.1	4.5	3.7
H. wuliangensis	23	12.0	11.0	12.5	14.3	13.7	14.1	14.0	13.8	12.7	12.5	12.8	12.5	11.6	11.7	12.9	12.6	12.6	0.5	11.4	11.4	11.4	11.2
H. wenshanensis	24	8.1	8.2	9.2	9.9	9.7	10.0	8.4	8.4	8.0	7.5	9.2	9.3	9.0	9.1	10.4	9.9	3.7	13.0	9.0	9.9	9.6	8.5	12.5
H. dotui	25	14.1	14.7	14.7	14.8	14.2	14.7	14.3	13.5	13.0	13.1	13.3	13.3	13.8	13.7	14.6	15.2	13.7	16.6	13.4	14.7	15.0	13.9	16.0	12.9