Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae) Author Wagner, Philipp 0575CE45-AB5C-4493-A8AD-DD399E18284A Allwetterzoo Münster, Sentruper Strasse 315, D- 48161 Münster, Germany & Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA wagner@allwetterzoo.de Author Ihlow, Flora 10DB565B-DA19-484E-AE50-ED29945658E8 Museum of Zoology Senckenberg, Dresden, Königsbrücker Landstrasse 159, D- 01109 Dresden, Germany Author Hartmann, Timo A911B13A-B364-424E-8579-AE23820F1DC1 ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany Author Flecks, Morris 974BF839-926B-4039-9617-135F5CEC2596 ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany Author Schmitz, Andreas 5B1146D2-B9A6-48E6-9E4E-EB1C8ED1AE7A Natural History Museum of Geneva, UREC-Herpetology & Ichthyology, Route de Malagnou 1, CH- 1208 Genève, Switzerland Author Böhme, Wolfgang FFAC2972-9F52-404B-BA9C-489C7793FF8D ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany text Bonn zoological Bulletin 2021 2021-05-07 70 1 141 171 journal article 3714 10.20363/BZB-2021.70.1.141 7399d769-249a-47c9-b099-719ecd66851c 5643781

Table 5. Morphological variation in the examined specimens of the Calotes mystaceus complex. Values are given are as mean ± standard deviation and minimum – maximum. Characters marked with an asterisk were significantly different between species in the AN(C)OVA.

	C. bachae C. geissleri sp. n. C. goetzi sp. n.		C. mystaceus		C. vindumbarbatus sp. n.
	females males females males females (n = 6) (n = 10) (n = 2) (n = 4) (n = 33)	males (n = 38)	females (n = 2)	males (n = 7)	females (n = 2)	males (n = 3)
SVL	80.5 ± 6.35 88.6 ± 14.06 103 ± 15.56 114.5 ± 7.72 101.33 ± 114.16 ± 87 ± 16.97 82.29 ± 17.58 60 ± 18.38 71–86 56–102 92–114 106–122 10.32 16.34 75–99 58–101 47–73 83–126 84–145					67 ± 14 51–77
EyeEar	4.82 ± 0.58 5.92 ± 1.21 6.84 ± 1.36 8.11 ± 1.25 6.01 ± 0.76 7.88 ± 1.64 5.61 ± 0.86 5.09 ± 1.18 3.51 ± 1.56 3.91–5.61 3.56–7.15 5.88–7.8 6.95–9.45 4.66–7.73 5.33–11.46 5–6.21 3.49–6.56 2.41–4.61					4.35 ± 0.79 3.44–4.89
HeadH	14.88 ± 1.85 16.2 ± 2.95 17.43 ± 3.3 20.75 ± 3.88 17.81 ± 2.13 12.19–17.35 11.7–19.55 15.1–19.76 17.45–26.35 13.74–22.38	21.85 ± 4.18 13.55– 31.79	15.14 ± 14.64 ± 3.57 10.05 ± 3.73 9.87–20.03 2.97 12.5–17.78 7.95–12.15			11.02 ± 2.07 8.64–12.33
HeadL	19.82 ± 1.94 21.97 ± 3.53 24.71 ± 4.65 28 ± 2.71 23.75 ± 2.15 16.92–21.5 14.71–25.72 21.42–28 24.89–31.07 20.14–28.39	27.18 ± 5.44 7.8–36	21.12 ± 20.4 ± 4.13 15 ± 4.49 3.85 14.59–24.73 11.83–18.18 18.4–23.85			16.28 ± 3.47 12.28–18.43
HeadW*	15.01 ± 1.04 16.89 ± 3.22 19.66 ± 2.62 24.87 ± 3.86 17.97 ± 2.16 23.84 ± 5.2 15.82 ± 15.79 ± 3.65 12.66 ± 14.12–16.93 11.21–20.54 17.81–21.52 20.81–30 14.92–22.73 12.75– 1.95 11.19–21.36 3.85 37.13 14.44–17.2 9.94–15.39					13.12 ± 2.59 10.19–15.12
Interorb*	8.96 ± 0.73 9.92 ± 1.67 14.16 ± 1.92 12.3 ± 1.99 11.12 ± 0.99 12.48 ± 10.39 ± 8.9 ± 1.6 7.16 ± 2.22 7.85–9.73 6.7–11.69 12.8–15.52 9.92–14.69 9.2–12.7 1.85 1.77 6.62–10.87 5.59–8.73 8.75–16.65 9.13–11.64					7.74 ± 1.43 6.08–8.57
JawW	13.92 ± 0.93 15.07 ± 2.5 17.77 ± 1.79 20.98 ± 1.53 16.54 ± 1.44 12.82–15.56 10.74–18.36 16.5–19.03 19.32–22.44 14.2–19.65	20.01 ± 4.06 12.03– 37.39	14.91 ± 14.09 ± 2.73 11.84 ± 1.53 10.44–17.6 3.34 13.83–16 9.47–14.2			11.89 ± 2.47 9.13–13.9
NarEye	5.35 ± 0.85 6.15 ± 1.04 7.31 ± 1.55 7.89 ± 0.69 6.86 ± 0.73 7.76 ± 1.31 6.62 ± 2.05 5.99 ± 1.43 4.23 ± 1.03 3.75–6.19 4.34–7.62 6.22–8.41 7.38–8.9 5.25–8.44 5.31–10.71 5.17–8.07 3.61–7.41 3.5–4.96					4.69 ± 0.86 3.72–5.33
SnEye	9.19 ± 1.26 10.08 ± 1.85 12.01 ± 2.23 12.44 ± 0.78 11.39 ± 1.13 12.97 ± 10.14 ± 9.54 ± 2.2 6.97 ± 1.82 7.58–10.56 6.32–11.96 10.43–13.59 11.71–13.5 8.74–14.22 2.19 2.67 6.44–11.72 5.69–8.26 8.64–17.54 8.26–12.03					7.69 ± 1.5 5.96–8.63
SnW	5.04 ± 0.62 5.53 ± 0.75 6.33 ± 0.49 6.92 ± 0.45 6.56 ± 1.52 7.78 ± 2.88 5.42 ± 0.59 5.37 ± 0.66 4.45 ± 0.69 4.06–5.83 3.94–6.56 5.98–6.68 6.49–7.48 5.11–14.39 5.1–18.43 5–5.84 4.48–6.3 3.96–4.93					4.64 ± 0.81 3.7–5.15
4FingL	10.04 ± 1.05 10.67 ± 1.47 12.35 ± 0.64 13.69 ± 0.46 12.14 ± 1.12 13.15 ± 10.27 ± 10.2 ± 1.74 7.45 ± 1.34 8.56–10.93 7.44–12.04 11.89–12.8 13.41–14.37 9.99–14.14 1.64 1.89 7.63–12.08 6.5–8.4 9.82–16.03 8.93–11.6					8.09 ± 1.41 6.46–8.91
4ToeL	13.38 ± 0.93 15.03 ± 1.95 16.22 ± 0.76 17.68 ± 0.71 15.12 ± 1.57 11.83–14.59 11.04–16.92 15.68–16.76 16.88–18.59 12.75–20.31	17.43 ± 14.91 ± 2.5 14.07 ± 2.92 10.46 ± 2.19 13.14– 10.31–18.19 2.64 13.71– 16.68 8.6–12.33 21.89				11.84 ± 1.4 10.25–12.86
CrusL	17.22 ± 1.66 18.46 ± 2.92 20.91 ± 3.43 23.41 ± 0.94 20.57 ± 2.25 14.58–18.55 12.65–21.57 18.49–23.34 22.39–24.22 17.11–27.25	23.31 ± 2.91 17.55– 28.98	17.35 ± 3.44 14.92– 19.78	17.08 ± 3.81 12.59 ± 11.83–21.83 4.19 9.62–15.55		13.89 ± 2.96 10.48–15.81
ForefL	13.35 ± 0.79 14.62 ± 2.22 15.9 ± 1.09 18.25 ± 0.4 15.95 ± 1.68 17.71 ± 13.36 ± 13.88 ± 2.75 10.39 ± 12.11–14.17 10.29–17.73 15.13–16.67 17.67–18.56 13.98–22.19 2.03 1.34 9.97–17.8 2.64 13.16–21.4 12.41–14.3 8.53–12.26					11.39 ± 1.73 9.39–12.43
HindfL	23.75 ± 1.94 25.69 ± 3.18 28.39 ± 2.74 30.88 ± 0.81 27.31 ± 2.22 20.34–25.3 19.75–28.7 26.46–30.33 29.94–31.58 23.74–33.62	30.82 ± 3.25 23.36– 37.41	25.34 ± 3.85 22.62– 28.07	24.22 ± 4.89 17.89 ± 4.4 17.24–29.07 14.78–21		19.95 ± 3.35 16.09–22.12
LoArmL	14.45 ± 1.51 15.49 ± 2.93 17.39 ± 2.81 20.09 ± 1.31 17.5 ± 2.22 19.78 ± 3.3 15.42 ± 11.95–15.75 9.6–18.64 15.4–19.38 18.48–21.57 13.64–24.14 13.64– 3.44 27.73 12.99– 17.85			13.98 ± 3.27 10.1 ± 3.68 8.99–17.38 7.5–12.7		11.12 ± 2.17 8.62–12.51
SnForeL	27.83 ± 3.43 30.3 ± 11.23 32 ± 2.83 42 ± 5.6 35.21 ± 4.54 23–32 2–39 30–34 37–50 29–45	40.17 ± 31.5 ± 6.36 29.29 ± 8.04 21 ± 5.66 7.61 27–36 20–41 17–25 27–59				22.61 ± 4.1 18–25.83
TailH	6.72 ± 0.97 9.74 ± 2.07 10 ± 2.67 14.56 ± 1.31 9.79 ± 1.44 5.12–8.06 6.02–12.29 8.11–11.89 13.44–16.43 7.46–12.69	14.41 ± 9.27 ± 3.52 9.04 ± 2.95 4.88 ± 1.92 3.26 6.78–11.76 5.18–13.07 3.52–6.23 6.6–21				6.91 ± 2.69 3.81–8.61