Integrative approach to resolve the Calotes mystaceus Duméril & Bibron, 1837 species complex (Squamata: Agamidae) Author Wagner, Philipp 0575CE45-AB5C-4493-A8AD-DD399E18284A Allwetterzoo Münster, Sentruper Strasse 315, D- 48161 Münster, Germany & Department of Biology and Center for Biodiversity and Ecosystem Stewardship, Villanova University, 800 Lancaster Avenue, Villanova, Pennsylvania 19085, USA wagner@allwetterzoo.de Author Ihlow, Flora 10DB565B-DA19-484E-AE50-ED29945658E8 Museum of Zoology Senckenberg, Dresden, Königsbrücker Landstrasse 159, D- 01109 Dresden, Germany Author Hartmann, Timo A911B13A-B364-424E-8579-AE23820F1DC1 ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany Author Flecks, Morris 974BF839-926B-4039-9617-135F5CEC2596 ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany Author Schmitz, Andreas 5B1146D2-B9A6-48E6-9E4E-EB1C8ED1AE7A Natural History Museum of Geneva, UREC-Herpetology & Ichthyology, Route de Malagnou 1, CH- 1208 Genève, Switzerland Author Böhme, Wolfgang FFAC2972-9F52-404B-BA9C-489C7793FF8D ZoologischesForschungsmuseum AlexanderKoenig, Adenauerallee 160, D- 53113 Bonn, Germany text Bonn zoological Bulletin 2021 2021-05-07 70 1 141 171 journal article 3714 10.20363/BZB-2021.70.1.141 7399d769-249a-47c9-b099-719ecd66851c 5643781

APPENDIX IV. Morphological variation in the type specimens of the Calotes mystaceus complex. Values given as mean ± standard deviation and minimum – maximum.

	C. bachae		C. geissleri sp. n.		C. goetzi sp. n.		C. mystaceus	C. vindumbarbatus sp. n.
	holotype	paratypes (n = 5)	holotype	paratypes (n = 4)	holotype	paratypes (n = 2)	holotype	holotype	paratypes (n = 4)
SVL	90	85 ± 8.22 74–97	110	108 ± 12.11 92–120	118	97.5 ± 9.19 91–104	80	77	61 ± 13.95 47–73
EyeEar	5.30	5.19 ± 1.2 3.91–7.15	6.95	7.57 ± 1.49 5.88–9.45	8.20	5.55 ± 0.80 4.99–6.12	4.68	4.89	3.79 ± 1.09 2.41–4.71
HeadH	14.88	14.44 ± 2.24 12.19–18.22	17.45	18.51 ± 2.29 15.10–19.85	26.81	17.55 ± 1.07 16.80–18.31	13.56	12.33	10.21 ± 2.23 7.95–12.15
HeadL	21.80	20.84 ± 3.26 16.92–25.72	24.89	26.36 ± 3.44 21.42–29.22	29.60	24.27 ± 1.32 23.33–25.20	19.72	18.43	15.11 ± 3.53 11.83–18.18
HeadW	17.04	15.57 ± 2.57 13.71–20.02	20.81	22.00 ± 3.15 17.81–25.11	25.35	18.34 ± 3.38 15.95–20.73	14.83	14.05	12.66 ± 3.00 9.94–15.39
Interorb	10.09	9.31 ± 1.45 7.85–11.69	9.92	13.22 ± 1.60 11.8–15.52	12.73	10.98 ± 1.00 10.27–11.69	8.20	8.56	7.24 ± 1.64 5.59–8.73
JawW	15.00	14.40 ± 2.06 12.62–17.94	19.32	19.50 ± 2.46 16.50–22.44	20.02	17.34 ± 1.63 16.19–18.50	13.73	12.63	11.68 ± 2.75 9.13–14.20
NarEye	6.16	5.54 ± 1.21 3.75–6.97	7.38	7.47 ± 0.92 6.22–8.41	7.95	6.51 ± 0.39 6.23–6.78	5.88	5.33	4.30 ± 0.81 3.50–5.03
SnEye	10.44	9.54 ± 1.46 7.58–11.54	11.71	12.14 ± 1.31 10.43–13.59	13.76	11.40 ± 0.51 11.04–11.76	9.90	8.63	7.10 ± 1.48 5.69–8.48
SnW	5.45	5.37 ± 0.92 4.06–6.56	6.49	6.59 ± 0.46 5.98–7.09	7.23	6.17 ± 0.49 5.82–6.52	4.98	5.15	4.41 ± 0.68 3.70–5.06
4FingL	11.79	10.28 ± 0.84 8.88–10.94	13.56	13.12 ± 1.04 11.89–14.37	13.30	12.12 ± 0.30 11.90–12.33	10.40	8.89	7.57 ± 1.27 6.46–8.91
4ToeL	15.08	13.47 ± 1.09 11.83–14.59	18.59	16.70 ± 0.75 15.68–17.47	16.62	14.58 ± 0.40 14.30–14.86	14.07	12.42	11.01 ± 1.96 8.60–12.86
CrusL	19.21	17.38 ± 2.07 14.58–19.83	22.83	22.11 ± 2.53 18.49–24.22	24.48	20.88 ± 1.33 19.94–21.82	16.42	15.81	12.75 ± 3.14 9.62–15.55
ForefL	13.74	13.59 ± 1.35 12.11–15.59	17.67	17.14 ± 1.57 15.13–18.44	17.43	15.00 ± 0.28 14.80–15.19	13.59	12.43	10.63 ± 1.96 8.53–12.35
HindfL	26.14	24.34 ± 2.45 20.34–26.97	31.58	29.70 ± 2.22 26.46–31.53	30.61	27.39 ± 0.33 27.16–27.62	24.22	21.64	18.5 ± 3.61 14.78–22.12
LoArmL	15.59	14.95 ± 2.51 11.95–18.64	19.73	18.46 ± 2.22 15.40–20.60	20.82	17.75 ± 0.88 17.12–18.37	13.41	12.23	10.33 ± 2.66 7.50–12.70
SnForeL	30	30.40 ± 4.67 25.00–37.00	40	35.50 ± 4.65 30.00–41.00	30	32.00 ± 1.41 31–33	24	24	21.46 ± 4.6 17.00–25.83
TailH	8.71	7.49 ± 2.02 5.12–10.35	13.44	12.09 ± 2.87 8.11–14.42	15.1	9.48 ± 1.67 8.30–10.66	9.33	8.61	5.47 ± 2.25 3.52–8.31
TailL	199	188.25 ± 21.53 157–203	201	203 ± 17.78 189–223	236	214 ± 0 214–214	164	NA	114 ± 26.08 86–140
TailW	8.44	7.41 ± 1.68 5.27–9.57	11.75	11.78 ± 1.08 10.17–12.35	13.38	9.82 ± 0.03 9.80–9.84	8.25	7.42	6.3 ± 2.58 3.67–9.1
TrunkL	42	37.60 ± 2.97 34–41	54	54.25 ± 4.57 49–59	54	47.00 ± 1.41 46–48	43	35	28.75 ± 4.99 24–34
UpArmL	14.73	13.66 ± 1.62 11.77–15.78	16.64	17.64 ± 1.97 15.98–20.09	17.13	15.77 ± 1.48 14.72–16.81	16.33	10.96	9.85 ± 1.57 8.44–11.79
UpLegL	20.24	18.51 ± 2.78 14.85–21.83	22.42	22.43 ± 2.46 19.23–25.17	23.26	19.91 ± 0.11 19.84–19.99	16.52	14.75	13.22 ± 2.19 10.88–15.32
CanthR	9	8.8 ± 0.84 8–10	6	6 ± 0 6–6	9	8.5 ± 0.71 8–9	6	5	5.25 ± 0.5 5–6
Eyelid	14	13.2 ± 0.45 13–14	12	12.25 ± 1.26 11–14	14	12.5 ± 0.71 12–13	12	12	11.75 ± 1.26 10–13
HeadSLn	17	15 ± 1.73 14–18	14	16 ± 1.83 14–18	14	16 ± 0 16–16	13	15	15.75 ± 1.5 14–17
HeadSTr	16	16.4 ± 1.34 15–18	16	16 ± 0.82 15–17	16	17.5 ± 0.71 17–18	15	16	16.25 ± 0.96 15–17
Inflab	11	10.2 ± 0.45 10–11	9	10.5 ± 1.29 9–12	10	10.5 ± 0.71 10–11	10	10	10.25 ± 0.5 10–11
	holotype	paratypes (n = 5)	holotype	paratypes (n = 4)	holotype	paratypes (n = 2)	holotype	holotype	paratypes (n = 4)
SnS	9	8.6 ± 0.89 8–10	7	6.75 ± 0.5 6–7	7	7.5 ± 0.71 7–8	7	7	6.5 ± 0.58 6–7
Suplab	10	10.2 ± 0.45 10–11	11	11 ± 0.82 10–12	9	10 ± 0 10–10	10	9	10.5 ± 0.58 10–11
4FingLm	21	20 ± 1.41 18–22	24	20.25 ± 2.06 18–23	19	20 ± 1.41 19–21	20	21	21 ± 0.82 20–22
4ToeLm	23	23.8 ± 1.3 22–25	28	25.25 ± 2.87 22–29	22	22.5 ± 0.71 22–23	25	26	26 ± 1.15 25–27
VertS	42	42 ± 4.18 35–46	44	46.5 ± 3.7 42–50	50	47.5 ± 4.95 44–51	41	40	46.25 ± 1.71 44–48
MidbS	48	45.8 ± 1.48 44–48	50	55.5 ± 5 50–62	52	52 ± 4.24 49–55	44	50	52.5 ± 1.91 50–54