A revision of the genus Sclerocoelus Marshall (Diptera: Sphaeroceridae)
Author
Kuwahara, Gregory K.
2E2D313F-7249-4F76-AF5D-615516840188
University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.
gkuwahar@gmail.com
Author
Marshall, Stephen A.
6C1C1C9E-365F-48BF-B8C7-31163EC5F3A7
University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.
samarsha@uoguelph.ca
Author
Paiero, Steven M.
B172670C-2159-4D6B-9542-53406F315DC7
University of Guelph Insect Collection and Insect Systematics Laboratory, School of Environmental Sciences, University of Guelph, Guelph, Ontario, N 1 G 2 W 1, Canada.
paieros@uoguelph.ca
text
European Journal of Taxonomy
2025
2025-03-04
979
1
277
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2803/12807
journal article
308980
10.5852/ejt.2025.979.2803
987b5fa2-ceec-45dd-bc15-a17fb7e3547d
2118-9773
14980852
F40A49A4-5DCC-491E-9D0A-7A3C2EC6D186
Table 1 (continued on next page). Character state matrix of Sclerocoelus Marshall, 1995, including Chespiritos Marshall, 2000, Paramosina Marshall, 2014, and Parasclerocoelus Marshall & Dong, 2008 as the outgroups. The plesiomorphic state is indicated by (0) and apomorphic state(s) by (1–5); multistate characters are treated as unordered.| Taxon | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | 11 | 12 | 13 | 14 | 15 | 16 | 17 | 18 |
|---|
| Chespiritos Marshall, 2000 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 |
|---|
| Paramosina Marshall, 2014 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| Parasclerocoelus Marshall & Dong, 2008 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
|---|
| Sclerocoelus aduncus sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. alpinus sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| S. altus sp. nov. | 0 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. andensis Marshall, 1997 | 1 | 1 | 0 | 0 | 1 | 4 | 1 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. argentinensis sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. azulensis sp. nov. | 0 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| S. binus sp. nov. | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 2 | 1 | 4 | 0 | 0 | 0 | 0 | 2 | 1 | 2 | 2 |
|---|
| S. brasilensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. bucki sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. caligarius sp. nov. | 1 | 1 | 1 | 0 | 2 | 4 | 1 | 2 | 1 | 4 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 2 |
|---|
| S. caribensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. chilensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. copiosus sp. nov. | 0 | 1 | 0 | 1 | 2 | 4 | 2 | 1 | 0 | 4 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 |
|---|
| S. costaricensis sp. nov. | 0 | 1 | 1 | 2 | 2 | 1 | 1 | 2 | 1 | 0 | 0 | 3 | 0 | 1 | 1 | 0 | 1 | 2 |
|---|
| S. cubus sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. dasysternum sp. nov. | 0 | 1 | 0 | 1 | 2 | 1 | 1 | 2 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 0 | 1 | 2 |
|---|
| S. dominicensis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 0 | 2 |
|---|
| S. dryadalis sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. elephas sp. nov. | 1 | 1 | 0 | 2 | 2 | 4 | 2 | 1 | 0 | 4 | 2 | 0 | 0 | 0 | 2 | 1 | 2 | 2 |
|---|
| S. espeletia sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 1 | 1 | 1 | 2 | 1 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. flavus sp. nov. | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 3 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. frigidifrons sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| S. galapagensis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 1 | 1 | 1 | 4 | 1 | 3 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. grandicercus sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 1 | 2 | 1 | 0 | 1 | 3 | 0 | 0 | 1 | 0 | 1 | 0 |
|---|
| S. hemorrhoidalis Marshall, 1997 | 1 | 1 | 0 | 0 | 2 | 4 | 2 | 1 | 1 | 4 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. inornatus sp. nov. | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. irregularis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 0 | 2 |
|---|
| S. latibarbus sp. nov. | 0 | 1 | 1 | 2 | 2 | 4 | 2 | 1 | 1 | 2 | 0 | 3 | 0 | 1 | 1 | 0 | 1 | 2 |
|---|
| S. lazulita sp. nov. | 0 | 1 | 0 | 0 | 2 | 2 | 0 | 0 | 0 | 2 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| S. limbus sp. nov. | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 |
|---|
| S. longibarbus sp. nov. | 0 | 1 | 1 | 1 | 2 | 4 | 2 | 1 | 1 | 0 | 0 | 2 | 1 | 0 | 2 | 0 | 0 | 0 |
|---|
| S. lutosus sp. nov. | 0 | 0 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 |
|---|
| S. mandibulum sp. nov. | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 5 | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 2 |
|---|
| S. masneri sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 2 | 1 | 1 | 4 | 0 | 4 | 0 | 0 | 1 | 0 | 2 | 2 |
|---|
| S. meridensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. nebulosus sp. nov. | 0 | 1 | 0 | 0 | 2 | 2 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. nitidistylus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 1 | 2 | 1 | 3 | 0 | 1 | 0 | 0 | 1 | 2 |
|---|
| S. ocellatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 1 | 3 | 1 | 4 | 0 | 0 | 2 | 0 | 1 | 2 |
|---|
| S. paranebulosus sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. pararegularis sp. nov. | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | ? | ? | ? | ? |
|---|
| S. parasordipes sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 0 | 5 | 0 | 1 | 0 | 0 | 2 | 0 | 2 | 1 |
|---|
| S. penai sp. nov. | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. plumiseta (Duda, 1925) | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 3 | 1 | 0 | 2 | 0 | 0 | 2 |
|---|
| S. punensis sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 0 | 1 | 2 | 1 | 2 | 0 | 0 | ? | ? | ? | ? |
|---|
| S. puyensis sp. nov. | 0 | 1 | 1 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 2 | 2 | 0 | 0 | 2 | 0 | 1 | 2 |
|---|
| S. rectangularis (Malloch, 1914) | 0 | 1 | 0 | 0 | 2 | 4 | 2 | 2 | 0 | 1 | 0 | 3 | 1 | 0 | 2 | 0 | 2 | 2 |
|---|
| S. recurvatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 2 | 2 | 2 | 1 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 3 | 0 |
|---|
| S. regularis (Malloch, 1914) | 0 | 1 | 0 | 3 | 1 | 1 | 1 | 1 | 1 | 3 | 0 | 2 | 1 | 2 | 1 | 0 | 2 | 2 |
|---|
| S. riparius sp. nov. | 0 | 1 | 0 | 0 | 1 | 3 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 |
|---|
| S. rostrum sp. nov. | 0 | 1 | 1 | 1 | 2 | 3 | 2 | 1 | 1 | 2 | 0 | 3 | 0 | 1 | ? | ? | ? | ? |
|---|
| S. sordipes (Adams, 1904) | 0 | 1 | 0 | 1 | 2 | 3 | 1 | 2 | 0 | 5 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 |
|---|
| S. subbrevipennis (Frey, 1954) | 1 | 1 | 0 | 0 | 2 | 4 | 1 | 1 | 1 | 4 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 |
|---|
| S. synorios sp. nov. | 0 | 1 | 1 | 1 | 2 | 4 | 2 | 1 | 1 | 1 | 1 | 3 | 0 | 1 | 2 | 0 | 0 | 2 |
|---|
| S. tantus sp. nov. | 1 | 1 | 0 | 2 | 2 | 4 | 2 | 2 | 0 | 2 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 |
|---|
| S. tridens sp. nov. | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 1 |
|---|
| S. turpis sp. nov. | 0 | 1 | 0 | 2 | 1 | 1 | 1 | 1 | 1 | 3 | 1 | 2 | 1 | 0 | ? | ? | ? | ? |
|---|
| S. vulgatus sp. nov. | 0 | 1 | 0 | 1 | 2 | 3 | 2 | 2 | 2 | 2 | 1 | 2 | 0 | 1 | 1 | 0 | 0 | 2 |
|---|
| S. xynos sp. nov. | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 |
|---|