A New Species of Helicops (Serpentes: Dipsadidae: Hydropsini) from Southeastern Brazil Author Costa, Henrique C. Author Santana, Diego J. Author Leal, Fernando Author Koroiva, Ricardo Author Garcia, Paulo C. A. text Herpetologica 2016 2016-06-30 72 2 157 166 http://dx.doi.org/10.1655/herpetologica-d-15-00059 journal article 231400 10.1655/HERPETOLOGICA-D-15-00059 c3c8c82e-cdab-46c5-9f80-f1ccd8fa0626 1938-5099 7712201

TABLE 4.—Descriptions of the color patterns seen in Helicops species.

Species	Dorsal color pattern	Ventral color pattern	Reference *
* (1) Boulenger 1893; (2) Amaral 1922; (3) Amaral 1938; (4) Rossman 1974; (5) Rossman 1975; (6) Rossman and Dixon 1975; (7) Rossman 1976; (8) Rossman and Abe 1979; (9) Rossman 2002a; (10) Rossman 2002b; (11) Bartlett and Bartlett 2003; (12) Frota 2005; (13) Yuki and Lema 2005; (14) Rossman 2010; (15) Kawashita-Ribeiro et al. 2013; (16) present study.
H. angulatus	Olive gray, tan or olive brown, with reddish brown to dark brown crossbands.	Cream to light tan, with or without extensions of dorsal crossbands, making a checkered	11, 16
pattern.
H. apiaka	Approximately 15 dark crossbands.	Creamy white with the extensions of large black dorsal crossbands.	15
H. carinicaudus	Uniformly dark-brown or black, sometimes with broad longitudinal stripes lighter	Cream with two (sometimes three) medial rows of black semilunar marks.	13, 16
than ground color.
H. danieli	Reddish-brown or olive-brown with 4–5 rows of alternating dark spots.	Cream with two medial rows of black semilunar marks, sometimes fused midventrally.	3, 9
H. gomesi	Olive-brown with irregular dark blotches.	Yellowish-brown or olive, with the extensions of dark dorsal blotches laterally.	2
H. hagmanni	Gray-brown with alternating light and dark blotches.	Irregularly checkered light/red and dark color.	5
H. infrataeniatus	Dark-brown with light-brown stripes.	Cream or red with 1–3 dark stripes or darkly checkered.	13, 16
H. leopardinus	Medium brown with dark brown or black blotches.	Checkerboard of black and red, banded black and red, or a combination of both.	11, 16
H. modestus	Dark brown with 2–3 narrow light brown stripes, or uniformly dark olive.	Cream, with or without faint brown flecks.	4, 15, 16
H. pastazae	Tan to gray brown with four or five rows of alternating dark spots, the paravertebral	Cream with a series of dark crossbands or alternating checks, the light ventral color	7
	spots large and often fused transversely.	extending onto several dorsal scale rows.
H. petersi	Dark olive brown with four or five rows of alternating dark spots.	Cream with a lateral series of dark checks.	7
H. polŋlepis	Brownish or olive gray, with blackish spots in five alternating rows.	Dark with pale spots.	1, 7
H. scalaris	Grayish tan with 3–5 rows of irregular dark blotches, the vertebral blotches larger	Cream medially with two rows of brown pigment forming either an irregular checkered	10
	than the laterals, and all 3 usually fused longitudinally on the neck to form short,	pattern or broad ventrolateral stripes, at least anteriorly.
broad stripes.
H. tapajonicus	Uniformly moss-green.	Checkered light and dark brown.	12
H. triƲittatus	Medium brown with five narrow longitudinal light stripes.	Cream with two medial rows of black semilunar markings.	14
H. ŋacu	Light to medium gray-brown, with four alternating rows of relatively small dark	Cream with an irregular series of large black checks or a median row of black crescents.	6, 8
spots.
H. nentur sp. nov. Uniformly dark-olive green, dark brown or dark gray.		Yellow (in life), cream or light brown (sometimes mottled with gray) with two medial	16
		rows of black semilunar markings.