The identity of the semiterrestrial crab Terrathelphusa kuchingensis (Nobili, 1901) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of four new species from southwestern Sarawak, Borneo, Malaysia Author Grinang, Jongkar Author Ng, Peter K. L. text Zootaxa 2015 3946 3 331 346 journal article 41512 10.11646/zootaxa.3946.3.2 d57d5862-5fc5-480c-84c9-3b598b597197 1175-5326 244741 24E2F41D-89BF-473F-ACE0-ED951BCB2699

TABLE 1. Differences between eight Terrathelphusa species from Borneo [* from Ng (1997)]

Last ambulatory merus	Relatively slender, length about 3.9 times width	Relatively slender, length about 3.9 times width	Relatively slender, length about 4.3 times width	Relatively slender, length about 3.8 times width	Relatively slender, length about 3.5 times width	Relatively slender, length about 4.4 times width	Relatively stout, length about 3.5 times width	Relatively stout length about 3.1 times width
Carpus of cheliped Sixth male abdominal segment	Surface rugose, without granules; inner angle with acutely triangular tooth Lateral margins gently concave	Surface rugose, without granules; inner angle with acutely triangular tooth Lateral margins gently concave	Surface rugose, without granules; inner angle with broadly triangular tooth Lateral margins gently concave	Surface rugose, without granules; inner angle with broadly triangular tooth Lateral margins gently concave	Surface with small, low to squamate granules; inner angle with acutely triangular tooth Lateral margins gently concave	Surface rugose, without granules; inner angle with thin and acutely triangular tooth Lateral margins gently concave	Surface with small, low to squamate granules; inner angle with acutely triangular tooth Lateral margins gently concave	Surface rugose, without granule; inner angle with broadly triangular tooth Lateral margin concave
First gonopods	Gently curving outwards; tip of terminal segment tapered; terminal segment, about 0.3 times length of subterminal segment	Gently curving outwards; tip of terminal segment tapered; terminal segment, about 0.3 times length of subterminal segment	Gently curving outwards; tip of terminal segment tapered; terminal segment, about 0.3 times length subterminal segment	Gently curving outwards; tip of terminal segment tapered; terminal segment, about 0.3 times length of subterminal segment	Gently curving outwards; tip of terminal segment flared; terminal segment about 0.3 times length of subterminal segment	Distinctly curving outwards; tip of terminal segment tapered; terminal segment, about 0.3 times length of subterminal segment	Strongly curving outwards; tip of terminal tapering to blunt tip; terminal segment 0.6-0.7 times length of subterminal segment	Strongly curving outwards; tip of terminal segment tapering to blunt tip; terminal segment about 0.5 times length of subterminal segment
Second gonopods	Distal segment relatively long, about 0.8 times length of basal segment	Distal segment relatively long, about 0.7 times length of basal segment	Distal segment relatively long, about 0.6 times length of basal segment	Distal segment relatively long, about 0.6 times length of basal segment	Distal segment relatively long, about 0.6 times length of basal segment	Distal segment relatively long, about 0.8 times length of basal segment	Distal segment relatively short, about 0.3 times length of basal segment	Distal segment relatively short, about 0.5 times length of basal segment
Live colour	Dark crimson carapace, reddish to purplish chelipeds and legs	Yellowish overall	Dark purple, orange to white chelipeds and legs	Grape purple carapace, white cream chelipeds and legs	No record	Golden yellow overall	Brown olive green overall	No record