Description of the first cryptobranch onchidoridid Onchimira cavifera gen. et sp. nov., and of three new species of the genera Adalaria Bergh, 1879 and Onchidoris Blainville, 1816 (Nudibranchia: Onchidorididae) from Kamchatka waters Author Martynov, Alexander Author Korshunova, Tatiana Author Sanamyan, Nadezhda Author Sanamyan, Karen text Zootaxa 2009 2159 1 43 journal article 40777 10.5281/zenodo.188931 2c4c696d-084b-4bb7-93c1-a39235211ea5 1175-5326 188931

TABLE 1. Comparison of the genera of the family Onchidorididae possessing dorsal gills

	Onchimira gen.nov.	Calycidoris Abraham, 1876	Acanthodoris Gray, 1850	Adalaria Bergh, 1879	Loy Martynov, 1994	Onchidoris Blainville, 1816
Notum consistency	Soft	Hard	Soft	Hard or soft	Soft	Hard or soft
Notum Rhinophoral sheaths Oral veil shape	Almost smooth or with low tubercles Raised, smooth, even edges Subtrapezoid, wide	Numerous spiniform tubercles No definite borders, except for several adjacent tubercles Two narrow elevations	Numerous spiniform tubercles (smooth in a single species) With raised crenulated edges Semicircular, wide, with two distinct lower lobes	Well defined wide spiculose tubercles No definite borders, except for several adjacent tubercles Semicircular, wide, entire in most species	Smooth Without definite borders and tubercles Two narrow separate triangular lobes	Wide spiculose tubercles in most species No definite borders, except for adjacent tubercles Semicircular, wide, entire
Gill shape and arrangement	Unipinnate, in complete circle	Unipinnate, in complete circle	Tripinnate, in a semi- circle	Uni- to tripinnate, semircle or almost complete circle	Unipinnate, placed at single point	Unipinnate, in a semicircle
Gill cavity (pocket) Gill retractibility Buccal pump Radular formula Labial cuticle	Present, fully contractable Fully retractable Sessile 7–9.1.1.1.9–7 Distinct rods	Present, similar to cryptobranch type but apparently incapable of full contraction Capable of partial retraction into gill cavity Sessile 3.1.0.1.3 Distinct rods	Absent Contractile Sessile 2–7.1.0.1.7–2 Distinct short irregularly denticulated rods	Absent Contractile Sessile or on short broad stalk 3–13.1.1–0.1.13–3 Indistinct polygonal elements	Present, of a special type?Contractile Sessile, low elevation of the anterior part of pharynx 7.1.1.0.1.1.7 Smooth	Absent Contractile Long stalked 2–0.1.0–1.1.0–2 Indistinctpolygonal elements
Stomach caecum	Absent	Present	Absent	Absent	Absent	Absent
Evertable part of vas deferens Oviduct pattern	Wide permanent penis with smaller inner evertable part Proximal oviduct enters female gland mass	Long narrow, forming penis while everted Proximal oviduct enters female gland mass	Long narrow, forming penis while everted Proximal oviduct enters female gland mass	Long narrow, forming penis while everted Proximal oviduct connects to allosperm receptacles or vagina	Wide short flattened permanent penis Proximal oviduct connects to allosperm receptacles	Long narrow, forming penis while everted Proximal oviduct connects to allosperm receptacles or vagina
Seminal receptacle pattern	Swollen area at terminal vagina; sessile	Long stalked; vaginal arrangement	Long stalked; vaginal arrangement	Short stalk at terminal part of the vagina	Swollen part of proximal oviduct	Long duct from terminal part of the vagina
Vagina	Short	Very long	Very long	Moderately long to short	Short	Short
Source	Present study	Abraham, 1876; Roginskaya, 1972; Martynov, 2006; Present study	Alder & Hancock, 1845- 1855; MacFarland, 1925, 1926; Thompson & Brown, 1984; Fahey & Valdés, 2005; Present study	Alder & Hancock, 1845- 1855; Bergh, 1879, 1880; Thompson & Brown, 1984; Millen, 1987, 2006; Martynov et al., 2006; Present study	Martynov, 1994a	Alder & Hancock, 1845- 1855; Schmekel & Portmann, 1982; Thompson & Brown, 1984; Millen, 1985; Martynov et al., 2006; Present study