Metabetaeus lapillicola, a new species of alpheid shrimp (Decapoda: Caridea) from Japan, inhabiting unusual pebble beaches Author Yamashita, Ryunosuke 0000-0003-3781-6663 School of Marine Resources and Environment, Tokyo University of Marine Science and Technology, 4 - 5 - 7 Konan, Tokyo, Minato 108 - 8477 Japan. ryamashita 0613 @ gmail. com; https: // orcid. org / 0000 - 0003 - 3781 - 6663 ryamashita0613@gmail.com Author Komai, Tomoyuki Natural History Museum and Institute, Chiba, 955 - 2 Aoba, Chiba, Chuo 260 - 8682, Japan Author Kon, Koetsu 0000-0003-1379-0702 Department of Ocean Sciences, Tokyo University of Marine Science and Technology, 4 - 5 - 7 Konan, Tokyo, Minato 108 - 8477 Japan kon @ kaiyodai. ac. jp; https: // orcid. org / 0000 - 0003 - 1379 - 0702 kon@kaiyodai.ac.jp text Zootaxa 2023 2023-03-29 5258 3 317 330 http://dx.doi.org/10.11646/zootaxa.5258.3.5 journal article 240356 10.11646/zootaxa.5258.3.5 af4f2d3b-196f-4413-b197-cc7c7962276a 1175-5326 7780429 E7459C8A-4B2E-45CA-B8ED-5137A4708F87

TABLE 2. Genetic divergence of the mitochondrial 16S rRNA gene among four species of Metabetaeus (10 sequences) based on Kimura 2-parameter model. The rate variation among sites was modeled with a gamma distribution (shape parameter = 1). There was a total of 386 bp in the final dataset.

	1	2	3	4	5	6	7	8	9
1. Metabetaeus lapillicola n. sp. (LC744376)
2. Metabetaeus lapillicola n. sp. (LC744377)	0.0028
3. Metabetaeus lapillicola n. sp. (LC744378)	0.0000	0.0026
4. Metabetaeus mcphersonae (KC879831)	0.1807	0.1964	0.1964
5. Metabetaeus lohena (KC879830)	0.1898	0.2110	0.2063	0.2256
6. Metabetaeus lohena (MZ661695)	0.1898	0.2110	0.2063	0.2256	0.0000
7. Metabetaeus minutus (KC879829)	0.1792	0.1906	0.1863	0.1960	0.1497	0.1497
8. Metabetaeus minutus (KC879828)	0.1579	0.1699	0.1658	0.1876	0.1303	0.1303	0.0244
9. Metabetaeus minutus (KC879827)	0.1784	0.1940	0.1897	0.1870	0.1254	0.1254	0.0571	0.0418
10. Metabetaeus minutus (MZ661696)	0.1621	0.1740	0.1699	0.1920	0.1303	0.1303	0.0188	0.0106	0.0418