The oldest Cenozoic ant fossil: † Tyrannomecia gen. nov. (Formicidae: Myrmeciinae) from the Palaeocene Menat Formation (France) Author Jouault, Corentin 0000-0002-3680-5172 Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier 75005, Paris, France & Univ. Rennes, CNRS, Géosciences Rennes, UMR 6118, F- 35000, Rennes, France & CNRS, UMR 5554 Institut des Sciences de l’Évolution de Montpellier, Place Eugène Bataillon, 34095, Montpellier, France jouaultc0@gmail.com Author Nel, André 0000-0002-4241-7651 Institut de Systématique, Évolution, Biodiversité (ISYEB) Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier 75005, Paris, France text Historical Biology 2021 2021-12-12 34 11 2241 2248 http://dx.doi.org/10.1080/08912963.2021.2010192 journal article 10.1080/08912963.2021.2010192 616f0513-90fe-4370-bb63-c7f70a876f26 1029-2381 5838068

Table 1. Fossil record of the subfamily Myrmeciinae

Genus	Species	Country	Age	Locality/Formation	Reference
Archimyrmex
	A. piatnitzkyi Viana and Haedo Rossi	Argentina	Lutetian	Pilcaniyeu road/Ventana Formation	Viana and Haedo Rossi 1957
1957
	A. rostratus Cockerill, 1923	USA	Bridgerian	Ute Trail, Roan Plateau/Green River	Cockerell 1923
				Formation
	A. smekali Rossi de Garcia 1983	Argentina	Lutetian	Confluencia/Ventana Formation	Rossi de Garcia 1983
	A. wedmannae Dlussky 2012	Germany	Lutetian/	Grube Messel Pit/Messel Formation	Dlussky 2012
			Ypresian
Avitomyrmex
	A. elongatus Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	A. mastax Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	A. systenus Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
Macabeemyrma
	M. ovata Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
Myrmeciites
	M. goliath Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	M. herculeanus Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	M. tabanifluviensis Archibald et al. 2006 Canada		Ypresian	Black Creek Road Locality	Archibald et al. 2006
				(Horsefly River)
Prionomyrmex
	P. gusakovi Radchenko and Perkovsky	Russia	Priabonian	Baltic amber/Yantarnyi,	Radchenko and Perkovsky
	2020			Kaliningrad Region	2020
	P. janzeni Baroni Urbani 2000	Russia	Priabonian	Baltic amber	Baroni Urbani 2000
	P. longiceps Mayr 1868	Baltic Sea	Priabonian	Baltic amber	Mayr 1868
		region
	P. wappleri Dlussky 2012	Germany	Oligocene	Rott	Dlussky 2012
Propalosoma
	P. gutierrezae Dlussky and Rasnitsyn	USA	Ypresian	Republic, locality B4131/Klondike	Dlussky and Rasnitsyn 1999
	1999			Mountain Formation
Tyrannomecia gen.
nov.
	T. inopinata sp. nov.	France	Selandian	Old quarry/Menat	This study
Ypresiomyrma
	Y. bartletti Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	Y. orbiculata Archibald et al. 2006	Canada	Ypresian	McAbee	Archibald et al. 2006
	Y. orientalis Dlussky et al. 2015	Russia	Priabonian	Bolshaya Svetlovodnaya, Biamo,	Dlussky et al. 2015
				layer 1
	Y. rebekkae Rust and Andersen 1999	Denmark	Ypresian	Stolleklint/Fur Formation	Rust and Andersen 1999