The very rare Japanese endemic diving beetle Japanolaccophilus niponensis (Kamiya, 1939), (Coleoptera: Dytiscidae, Laccophilinae): larval morphology and phylogenetic comparison with other known Laccophilini Author Alarie, Yves School of Natural Sciences, Laurentian University, Ramsey Lake Road, Sudbury, Ontario, CANADA. Author Watanabe, Kohei Ishikawa Insect Museum, Inu- 3, Yawata-machi, Hakusan-shi, 920 - 2113 JAPAN. Author Michat, Mariano C. 0000-0002-1962-7976 University of Buenos Aires, Faculty of Exact and Natural Sciences, Department of Biodiversity and Experimental and Applied Biology (IBBEA), Buenos Aires, ARGENTINA. marianoide @ gmail. com; https: // orcid. org / 0000 - 0002 - 1962 - 7976 marianoide@gmail.com text Zootaxa 2023 2023-05-15 5285 1 116 132 http://dx.doi.org/10.11646/zootaxa.5285.1.4 journal article 10.11646/zootaxa.5285.1.4 1175-5326 7935858 0703FF00-A0C3-4A18-8437-4E25FE1BA047

TABLE 4. Measurements and ratios for the instar III of selected genera of Laccophilinae 1: Africophilus Guignot, 1947 (AFRI); Agabetes Crotch, 1873 (AGAB); Australphilus Watts, 1978 (AUST); Japanolaccophilus Satô, 1972 (JAPA); Laccomimus Toledo and Michat, 2015 (LACM); Laccophilus Leach, 1815 (LACP); Neptosternus Sharp, 1882 (NEPT); Philodytes J. Balfour-Browne, 1938 (PHYL); n, number of species (number of specimens measured).

Measure	AGAB (n = 1(2))	AFRI (n = 1(1))	AUST (n = 2(6))	JAPA (n = 1(4))	LACM (n = 1(3))	LACP (n = 10(35))	NEPT (n = 2(10))	PHYL (n = 1(2))
1, Alarie et al. 2000, 2002; Miller et al. 2005; Michat 2008; Michat & Toledo 2015; Watanabe & Okada 2023.
HL (mm)	1.26–1.37	0.59	0.78–0.82	0.84–0.89	0.53–0.54	0.95–1.43	0.82–0.98	1.67–1.77
HW (mm)	1.24–1.27	0.61	0.71–0.81	0.77–0.80	0.47–0.48	0.77–1.29	0.69–0.84	1.38–1.45
FRL (mm)	0.56–0.59	0.34	0.38–0.46	0.45–0.47	0.27–0.30	0.50–0.81	0.38–0.47	0.79–0.82
OCW (mm)	0.72–0.73	-----	0.33–0.35	0.38–0.44	0.22–0.25	0.39–0.72	0.30–0.38	0.80–0.90
HL/HW	1.02–1.08	0.97	1.00–1.09	1.09–1.11	1.13–1.17	1.06–1.30	1.13–1.21	1.16–1.29
HW/OCW	1.69–1.75	-----	2.15–2.50	1.76–1.81	1.92–2.18	1.64–2.08	2.13–2.33	1.53–1.80
COL/HL	0.55–0.57	0.43	0.43–0.52	0.47–0.50	0.45–0.49	0.40–0.54	0.49–0.54	0.53–0.54
A/HW	0.54–0.59	0.45	0.68–0.71	0.72–0.76	0.43–0.46	0.63–0.75	0.65–0.73	0.72–0.75
A3/A1	1.02–1.24	2.05	1.09–1.20	1.25–1.44	2.14–2.50	0.93–1.26	0.83–1.02	0.87–0.94
A3/A2	1.02–1.13	0.72	1.14–1.24	1.10–1.22	0.93–1.00	1.04–1.56	0.88–1.08	1.03–1.07
A4/A1	0.21–0.26	1.40	0.29–0.37	0.35–0.42	0.86–1.00	0.17–0.29	0.27–0.39	0.13–0.14
A4/A3	0.20–0.21	0.68	0.27–0.35	0.27–0.31	0.40–0.43	0.17–0.29	0.32–0.46	0.15–0.16
A3’/A4	0.85–0.91	0.34	0.72–1.00	0.58–0.63	0.83–1.00	0.67–0.96	0.83–1.10	0.66–0.70
MNL/MNW	3.08–3.23	2.27	2.93–3.30	2.78–2.94	2.81–3.07	2.66–3.38	3.13–3.51	2.89–2.91
MNL/HL	0.52–0.53	0.43	0.47–0.51	0.48–0.50	0.41–0.42	0.43–0.54	0.45–0.54	0.53–0.55
A/MP	1.18–1.24	1.27	0.99–1.05	1.01–1.08	1.14–1.22	1.07–1.23	1.03–1.07	1.18–1.21
MP2/MP1	1.21–1.25	1.91	1.06–1.19	1.13–1.22	2.43–2.57	0.92–1.14	0.89–0.97	0.92
MP2/MP3	1.12–1.13	1.55	0.96–1.03	0.84–0.87	1.55–1.64	0.90–1.25	0.98–1.18	1.24–1.30
GA/MP1	0.36–0.37	0.53	0.14–0.20	0.27–0.36	0.43	0.28–0.44	0.10–0.14	0.33
PPF/MP1	0.24–0.29	0.66	0.19–0.25	0.25–0.32	0.71	0.18–0.28	0.38–0.49	0.26–0.29
MP/LP	2.86	2.03	2.82–3.10	2.28–2.47	2.77–3.00	2.28–2.89	2.32–2.69	2.64–2.75
LP2/LP1	2.25	1.70	1.15–1.29	1.24–1.52	2.00–2.25	0.95–1.50	0.89–1.10	0.92
L1 (mm)	2.66–2.70	1.09	1.53–1.82	1.82–1.90	0.97–0.99	2.15–3.12	1.51–1.91	3.54–3.55
L2 (mm)	3.05–3.20	1.23	1.77–2.13	2.08–2.19	1.03–1.07	2.52–3.81	1.68–2.15	4.22–4.35
L3 (mm)	3.51–3.83	1.41	1.95–2.41	2.50–2.58	1.24–1.30	2.87–4.50	2.03–2.58	4.87–4.88
L2/L1	1.15–1.18	1.13	1.16–1.19	1.13–1.15	1.06–1.08	1.14–1.23	1.10–1.15	1.19–1.23
L3/L1	1.32–1.42	1.30	1.24–1.36	1.35–1.38	1.28–1.32	1.35–1.45	1.33–1.39	1.37–1.38
L3/L2	1.15–1.20	1.15	1.07–1.16	1.18–1.22	1.19–1.22	1.16–1.21	1.16–1.22	1.12–1.16
L3/HW	2.83–3.02	2.30	2.74–3.01	3.21–3.27	2.60–2.74	3.18–3.84	2.77–3.20	3.37–3.54
L3(CO/FE)	0.95–1.01	1.25	0.93–0.95	0.92–0.94	0.94–0.97	0.91–1.10	0.95–1.03	0.95–0.99
L3(TI/FE)	0.65–0.66	0.60	0.70–0.73	0.78–0.81	0.56–0.60	0.69–0.88	0.76–0.80	0.79–0.80
L3(TA/FE)	0.76–0.77	0.51	0.92–1.00	0.82–0.89	0.69–0.73	0.77–1.04	0.77–0.94	0.85–0.88
L3(CL/TA)	0.28–0.31	0.27	0.24–0.28	0.28–0.30	0.35–0.37	0.23–0.35	0.18–0.28	0.23–0.25
LAS (mm)	1.80–1.95	0.69	0.37–0.44	0.43–0.45	0.61–0.65	0.58–1.03	0.29–0.40	1.29–1.32
LAS/HW	1.45–1.54	1.13	0.53–0.56	0.54–0.58	1.29–1.38	0.64–0.93	0.41–0.48	0.91–0.94
U (mm)	0.20–0.22	0.28	0.88–0.98	0.83–0.86	0.14–0.16	1.40–2.40	0.72–1.13	2.54
U1 (mm)	0.20–0.22	0.13	0.84–0.98	0.75–0.79	0.08–0.09	1.39–2.37	0.71–1.11	2.54
U1/U2	-----	0.89	15.8–21.1	9.52–12.09	1.21–1.64	28.7–100.2	41.6–56.7	-----
U1/HW	0.16–0.17	0.22	1.14–1.22	0.97–0.99	0.18–0.20	1.42–2.30	0.97–1.31	1.76
U1/LAS	0.11–0.12	0.19	2.09–2.30	1.69–1.82	0.13–0.15	1.54–2.98	2.36–3.13	1.93