Begonia (sect. Ruizopavonia) De Candolle 1859 Author Moonlight, P. W. Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, Scotland, UK. E-mail: pmoonlight @ rbge. ac. uk. pmoonlight@rbge.ac.uk. Author Pérez, Á. J. Herbario QCA, Department of Biological Sciences, Pontificia Universidad Católica del Ecuador, Apartado Postal 17 - 01 - 2184, Quito, Ecuador. text Edinburgh Journal of Botany 2022 2022-08-18 79 401 1 9 http://dx.doi.org/10.5281/zenodo.8389628 journal article 285341 10.24823/EJB.2022.401 b6f8a384-203f-46e3-9af1-4e6cc7ccd440 1747-0036 10518476

Table. Distinguishing characters for northern Andean members of Begonia sect. Ruizopavonia with four tepals on the staminate flowers and elongate inflorescence internodesa

Character	B. pseudodendro n	B. consobrina	B. cuatrecasasiana	B. cymbalifera	B. dugandiana
a Key distinguishing characters are in bold italic text.
Distribution	Ecuador and Colombia, western slopes of the Andes, Nariño and Carchi	Ecuador, eastern slopes of the Andes, Sucumbios to Azuay	Colombia, western slopes of the western cordillera, Valle del Cauca to Antioquia	Ecuador and Colombia, eastern slopes of the Andes, Putumayo to Sucumbios	Colombia, both slopes of the Western cordillera, Antioquia to Tolima
Habit	Terrestrial herb to 10 m	Terrestrial herb to 1.5 m	Terrestrial or rarely epiphytic herb to 50 cm	Terrestrial herb to 1.5 m	Terrestrial herb to 1 m
Leaves	Straight, oblanceolate to elliptic	Transversely ovate	Straight, oblanceolate to elliptic	Straight, oblanceolate to ovate	Straight to transversely ovate
Venation	Pinnate, 7–9 secondary veins on the broad side of the lamina, 5–7 on the narrow side	Palmate-pinnate, 9–12 secondary veins on the broad side of the lamina, 7–9 on the narrow side	Pinnate, 6–8 secondary veins on the broad side of the lamina, 6 or 7 on the narrow side	Pinnate, 8–10 secondary veins on the broad side of the lamina, 6–8 on the narrow side	Palmate-pinnate 6–8 secondary veins on the broad side of the lamina, 4–6 on the narrow side
Outer two tepals on the staminate flowers	Ovate, apex rounded, 3–6 × 2–4 mm, white to pink, glabrous	Ovate, apex rounded, 4–5 × 4–5 mm, white to pink, glabrous	Ovate, apex long-acuminate, 20–28 × 12–15 mm, orange, pilose outside	Ovate, apex rounded, c.5 × 9 mm, white, glabrous	Ovate, rounded, c.5.5 × 5 mm, white, glabrous
Inner two tepals on the staminate flowers	Oblanceolate, c.4 × 1.5–2 mm, white to pink, glabrous	Oblanceolate to oblong, c.3 × 1 mm, white, glabrous	Oblong, 13–18 × 3.5–4 mm, orange, glabrous	Oblong, c.5 × 2 mm, white, glabrous	Oblong, c.3.5 × 1.5 mm, white, glabrous
Stamens	c.20	Numerous (> 25)	11–14	Numerous (> 25)	c.20
Bracteoles	Lacking	3, persistent	2, persistent	2 or 3, persistent	3, persistent
Tepals on pistillate flowers	2, 4–12 × 3–10 mm, white to pink, glabrous	3, the largest two c.8 × 8 mm, white to pink, glabrous	3, the largest two c.25 × 18 mm, orange, pilose outside	3, the largest two c.5 × 7 mm, white, glabrous	Unknown
Styles	Bifid	Multifid	Bifid	Multifid	Unknown