Including fossils in phylogeny: a glimpse into the evolution of the superfamily Evanioidea (Hymenoptera: Apocrita) under tip-dating and the fossilized birth-death process Author Jouault, Corentin Author Maréchal, Arthur Author Condamine, Fabien L. Author Wang, Bo Author Nel, André Author Legendre, Frédéric Author Perrichot, Vincent text Zoological Journal of the Linnean Society 2022 2021-07-13 194 1396 1423 https://doi.org/10.1093/zoolinnean/zlab034 journal article 56000 10.1093/zoolinnean/zlab034 ad63540f-94b2-4f6e-bb22-cdba0406f40c 0024-4082 6458537 9E9E6EC8-929E8-488A-AF43-0619A1820A7A

Table 1. Diversity and distribution of evanioid fossil genera

Families/subfamilies/genera	Species	Distribution	Period	References
Praeaulacidae
Cretocleistogastrinae
Cretocleistogaster Rasnitsyn, 1975	3	Russia	Late Cretaceous	Rasnitsyn (1975)
Miniwestratia Rasnitsyn, 1990	1	Mongolia	Late Cretaceous	Rasnitsyn (1990a)
Nanowestratia Rasnitsyn, 1990	1	Russia	Late Cretaceous	Rasnitsyn (1990a)
Sinowestratia Zhang & Zhang, 2000	1	China	Late Cretaceous	Zhang & Zhang (2000)
Westratia Rasnitsyn, 1990	3	Russia	Late Cretaceous	Rasnitsyn (1990a, b)
	4	Mongolia	Late Cretaceous	Rasnitsyn (1990a)
	1	Australia	Late Cretaceous	Jell & Duncan (1986)
Praeaulacinae
Archaulacus Li, Shih & Ren, 2014	1	China	Middle Jurassic	Li et al. (2014a)
Aulacogastrinus Rasnitsyn, 1972	1	Kazakhstan	Late Jurassic	Rasnitsyn (1972, 1983)
	3	China	Middle Jurassic	Zhang & Rasnitsyn (2008)
Eonevania Rasnitsyn & Zhang, 2010	1	China	Middle Jurassic	Rasnitsyn & Zhang (2010)
Eosaulacus Zhang & Rasnitsyn, 2008	1	China	Middle Jurassic	Zhang & Rasnitsyn (2008)
Evanigaster Rasnitsyn, 1972	1	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
Evaniops Rasnitsyn, 1972	1	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
Gulgonga Oberprieler, Rasnitsyn & Brothers, 20121		Australia	Late Jurassic	Oberprieler et al. (2012)
Habraulacus Li, Rasnitsyn, Shih & Ren, 2015	1	Myanmar	Late Cretaceous	Li et al. (2015)
Nevania Zhang & Rasnitsyn, 2007	8	China	Middle Jurassic	Zhang & Rasnitsyn (2007); Li et al. (2014b)
	1	Kazakhstan	Late Jurassic	Zhang & Rasnitsyn (2008)
Praeaulacinus Rasnitsyn, 1972	3	Kazakhstan	Late Jurassic	Rasnitsyn (1972, 1973)
Praeaulacites Rasnitsyn, 1972	5	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
Praeaulacon Rasnitsyn, 1972	4	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
Praeaulacops Rasnitsyn, 1972	1	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
Praeaulacus Rasnitsyn, 1972	6	Kazakhstan	Late Jurassic	Rasnitsyn (1972)
	2	Mongolia	Late Jurassic	Rasnitsyn (2008)
	9	China	Middle Jurassic	Zhang & Rasnitsyn (2008); Li et al. (2014a,
				2018); Li & Shih (2015)
Paleosyncrasis Poinar, 2019	1	Myanmar	Late Cretaceous	Poinar (2019)
Rasnitsevania Jouault, Nel & Perrichot, 2020	1	Myanmar	Late Cretaceous	Jouault et al. (2020)
Andreneliidae
Andrenelia Rasnitsyn & Martínez-Delclòs, 2000	1	Spain	Early Cretaceous	Rasnitsyn & Martínez-Delclòs (2000)
Anomopterellidae
Anomopterella Rasnitsyn, 1975	1	Kazakhstan	Middle Jurassic	Rasnitsyn (1975)
	1	Mongolia	Late Jurassic	Rasnitsyn (2008)
	8	China	Middle Jurassic	Zhang & Rasnitsyn (2008); Li et al. (2013b,
				2014b)
Choristopterella Li et al., 2013	1	Kazakhstan	Middle Jurassic	Rasnitsyn (1975)
Synaphopterella Li et al., 2013	1	China	Middle Jurassic	Li et al. (2013a)
Baissidae
Baissa Rasnitsyn, 1975	2	Russia	Early Cretaceous	Rasnitsyn (1975, 1991)
	1	Mongolia	Early Cretaceous	Rasnitsyn (1991)
Electrobaissa Engel, 2013	1	USA	Late Cretaceous	Engel (2013)
Heterobaissa Li et al., 2018	1	China	Early Cretaceous	Li et al. (2018)
Humiryssus Lin, 1980	5	China	Early Cretaceous	Lin (1980); Hong & Wang (1990); Zhang &
				Rasnitsyn (2004)
	1	UK	Early Cretaceous	Rasnitsyn et al. (1998)
Manlaya Rasnitsyn, 1980	3	China	Early Cretaceous	Ren (1995); Li et al. (2019)
	8	Mongolia	Early Cretaceous	Rasnitsyn (1980a, 1986)
	2	Russia	Early Cretaceous	Rasnitsyn (1980b)
	2	Spain	Early Cretaceous	Rasnitsyn & Martínez-Delclòs (2000)
	3	UK	Early Cretaceous	Rasnitsyn et al. (1998)
Mesepipolaea Zhang & Rasnitsyn, 2004	2	China	Early Cretaceous	Zhang & Rasnitsyn (2004); Li et al. (2019)
Tillywhimia Rasnitsyn & Jarzembowski, 1998	2	UK	Early Cretaceous	Rasnitsyn et al. (1998)
Othniodellithidae
Keratodellitha Jouault, Maréchal, Wang &	3	Myanmar	Late Cretaceous	This study
Perrichot gen. nov.
	1	Myanmar	Early Cretaceous	This study
Othniodellitha Engel & Huang, 2016	1	Myanmar	Late Cretaceous	Engel et al. (2016a)
Xenodellitha Engel, 2017	1	Myanmar	Late Cretaceous	Engel (2017)
Aulacidae
Paleoaulacus Jouault & Nel, 2021	1	Myanmar	Late Cretaceous	Jouault & Nel (2021)
Vectevania Cockerell, 1922	1	UK	Eocene	Cockerell (1922)
Aulacinae
Aulacus Jurine, 1807	1	France	Eocene	Nel et al. (2004)
	2	Baltic amber	Eocene	Brues (1933)
	1	USA	Oligocene	Brues (1910)
Electrofeonia Jouault et al., 2020	1	Myanmar	Late Cretaceous	Jouault et al. (2020)
Electrofoenops Engel, 2017	3	Myanmar	Late Cretaceous	Engel (2017); Turrisi & Ellenberger (2019)
Electrofoenus Cockerell, 1917	1	Myanmar	Late Cretaceous	Cockerell (1917)
Exilaulacus Li et al., 2018	2	Myanmar	Late Cretaceous	Li et al. (2018)
Pristaulacus Kieffer, 1900	3	USA	Eocene	Brues (1910); Cockerell (1916)
	3	Baltic amber	Eocene	Brues (1933); Jennings & Krogmann (2009)
Hyptiogastritinae
Archeofoenus Engel, 2017	2	Myanmar	Late Cretaceous	Engel (2017); Turrisi & Ellenberger (2019)
Hyptiogastrites Cockerell, 1917	1	Myanmar	Late Cretaceous	Cockerell (1917)
Protofoenus Cockerell, 1917	1	Myanmar	Late Cretaceous	Cockerell (1917)
Gasteruptiidae
Hypselogastriinae
Hypselogastrion Engel & Wang, 2016	1	Myanmar	Late Cretaceous	Engel & Wang (2016a)
Kotujellitinae
Kotujellites Rasnitsyn, 1975	1	Russia	Late Cretaceous	Rasnitsyn (1975)
Kotujisca Rasnitsyn, 1991	1	Mongolia	Late Cretaceous	Rasnitsyn (1991)
Evaniidae
Botsvania Rasnitsyn & Brothers, 2007	1	South Africa	Late Cretaceous	Rasnitsyn & Brothers (2007)
Brachygaster Leach, 1815	1	Dominican Republic	Miocene	Nel et al. (2002)
Burmaevania Shih et al., 2019	2	Myanmar	Late Cretaceous	Shih et al. (2019)
Cretevania Rasnitsyn, 1975	2	Russia	Late Cretaceous	Rasnitsyn (1975)
	1	Mongolia	Early Cretaceous	Rasnitsyn (1991)
	1	UK	Early Cretaceous	Rasnitsyn et al. (1998)
	5	Spain	Early Cretaceous	Peñalver et al. (2010); Pérez-De La Fuente
				et al. (2012)
	2	Myanmar	Late Cretaceous	Jennings et al. (2013); Li et al. (2018)
	4	China	Early Cretaceous	Zhang & Zhang (2000); Zhang et al. (2007);
				Li et al. (2014c, 2018)
	1	Jordan	Early Cretaceous	Kaddumi (2007)
	1	Lebanon	Early Cretaceous	Deans (2004)
Curtevania Li et al., 2018	1	Myanmar	Late Cretaceous	Li et al. (2018)
Eoevania Nel et al., 2002	1	France	Eocene	Nel et al. (2002)
Evaniella Bradley, 1905	1	Baltic amber	Eocene	Sawoniewicz & Kupryjanowicz (2003)
	2	Dominican Republic	Miocene	Poinar (2020)
Grimaldivania Basibuyuk et al., 2000	2	USA	Late Cretaceous	Basibuyuk et al. (2000)
Hyptia Illiger, 1807	2	Mexico	Miocene	Jennings et al. (2012); Poinar (2020)
	1	Baltic amber	Eocene	Jennings et al. (2013)
Iberoevania Peñalver et al., 2010	1	Spain	Early Cretaceous	Peñalver et al. (2010)
Lebanevania Basibuyuk & Rasnitsyn, 2002	1	Lebanon	Early Cretaceous	Basibuyuk et al. (2002)
Mesevania Basibuyuk & Rasnitsyn, 2000	1	Myanmar	Late Cretaceous	Basibuyuk et al. (2000)