First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan Author Yamana, Yusuke Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan Author Hirashima, Kentarou Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan Author Sato, Masanori Graduate School of Science and Engineering, Kagoshima University, Kagoshima 890 - 0065, Japan & Present address: 5233 Hiramatsu, Aira, Kagoshima 899 - 5652, Japan Author Yamamori, Luna Seto Marine biological laboratory (SMBL) of Kyoto University, 459, Shirahama, Wakayama 649 - 2211, Japan text Zootaxa 2025 2025-01-13 5569 1 1 54 https://doi.org/10.11646/zootaxa.5569.1.1 journal article 307817 10.11646/zootaxa.5569.1.1 e18dbbc9-ecbb-4d21-8b14-198f588ed3dd 1175-5326 14703528 96194340-054D-49C0-BD89-D47899BCAF14

TABLE 15-2. Comparison of the ossicles morphologies of previously nominated six species and four new species of the genus Patinapta Heding, 1928.

Specific name	Source		Tentacle		Longitudinal muscle						Body wall
*Length: Measurements denoted in original description or values in the present study based on holotype specimens or largest specimen (μm) a–i: (see, Materials and Methods) Indicators for ossicles shape, and one parameter (Ntp, number s teethed perforation) in original description (calculated as average values from illustrations for ossicles, extracted from different body parts by each author) and average values in the present study (middle body parts).
		Rod ossicles			Granule ossicles			Anchor plate ossicles				Anchor ossicles
Gen Patinapta	Author, Year	Length* RP a		RC b	Length*	GP c GCO d		Length*	APS e	Ntp f	APT g	Length*	ASW h	AEW i
P. crosslandii	Heding, 1929	Granules with both distal perforations			Often granules with a distal perforation			80–115	2.2	0	73.8	135	16.3	52.6
P. deformis sp. nov.	Present study	52–85	14.7	7.6	23–36	56.2	61.1	98–128	7.0	3.7	12.5	144–178	20.2	46.5
P. dumasi	Cherbonnier, 1954	-	-	-	Mostly O-shape, fewly C-shape granules			-	-	6–8	-	Both arms sharply turning to basal direction.
P. edentatus sp. nov.	Present study	Mostly elongated C-shape granules.			21–50	54.9	0	85–123	1.3	0	0	140–188	25.3	48.4
P. laevis	Bedford, 1899	Needle-like liner and narrow shape rods			Mostly two perforated oval granules.			83; 88	3.7	0	0	122; 144	11.3	61.6
P. neglectus sp. nov.	Present study	62–85	14.7	9.3	21–50	54.9	46.2	85–123	0–2.5	0–2	3.5	140–188	25.3	48.4
P. ooplax	von Marenzeller, 1882	52–87	13.1 11.2		28–40	44.3	50.0	94–109	0	7	22.6	113–119	21.9	55
P. ooplax	Present study	69–97	9.8	8.9	27–59	47.4	72.9	93–117	2.0	7.5	34.5	144–164	21.9	52.9
P. parvaspiculus sp. nov.	Present study	53–102 10.7		6.4	21–34	47.3	57.5	75–98	0–0.7	0–2	6.6	144–173	24.9	51.3
P. taiwaniensis	Chao et al. 1988	37–90	10.9	7.9	20–60	31.0	18.2	80–105	0.3	4.3	20.4	155–160	12.3	42.6
P. taiwaniensis	Present study	69–104 11.7		9.3	20–37	47.9	75.7	83–115	0.4	3.8	19.8	138–159	22.6	52.9
P. vaughani	Cherbonnier, 1953	63.5	7.0	5.5	Rod-like granules often with distal perforations			Most Anchor plates with four large teethless-perforations in distal area.				87–108	16.2	52.4