First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan Author Yamana, Yusuke Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan Author Hirashima, Kentarou Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan Author Sato, Masanori Graduate School of Science and Engineering, Kagoshima University, Kagoshima 890 - 0065, Japan & Present address: 5233 Hiramatsu, Aira, Kagoshima 899 - 5652, Japan Author Yamamori, Luna Seto Marine biological laboratory (SMBL) of Kyoto University, 459, Shirahama, Wakayama 649 - 2211, Japan text Zootaxa 2025 2025-01-13 5569 1 1 54 https://doi.org/10.11646/zootaxa.5569.1.1 journal article 307817 10.11646/zootaxa.5569.1.1 e18dbbc9-ecbb-4d21-8b14-198f588ed3dd 1175-5326 14703528 96194340-054D-49C0-BD89-D47899BCAF14

TABLE 9. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta parvaspiculus sp. nov.

Body parts	(Ossicle-type)	2015-INV-107		2023-INV-157		2024-INV-156
a–I, (see, Materials and Methods) Indicators for ossicle shapes, and one parameter (Ntp, number of teethed perforations).
Tentacle	Rods (n)	9		15		10
Length	(Range)	62.0±5.6	(55–74)	75.0±1.4	(53–102)	56.1±10.5	(45–79)
Breadth	(Range)	8.1±0.9	(7–10)	7.8±1.4	(6–11)	7.8±2.3	(5–12)
RP a (%)	(Range)	13.2±2.1	(10.8–16.9)	10.7±2.5	(7.1–16.9)	14.1±4.1	(8.2–20.7)
RC b	(Range)	8.0±1.7	(5.3–10.2)	6.4±2.3	(2.7–11.7)	13.4±6.2	(6.6–25.9)
Ventral side
L-Muscle	Granules (n)	12		11		12
Length	(Range)	28.4±4.3	(23–36)	27.8±2.3	(24–32)	29.6±4.2	(21–34)
Width	(Range)	16.0± 2.1	(10–21)	12.7±2.1	(10–18)	16.5±5.9	(12–33)
GP c	(Range)	50.7±15.5	(30.6–80.8)	45.9±7.8	(35.5–64.3)	33.3±0.5	(35.3–106)
GCO d (%)	(Ratio)	50.0	(O6: C6)	36.4	(O4: C7)	56.7	(O4: C8)
Body wall	Anchor plates (n)	5		7		9
Length	(Range)	87.6±7.6	(75–94)	90.9±5.6	(80–97)	93.0±3.5	(86–97)
Width	(Range)	49.0±5.8	(39–54)	57.3±4.9	(53–64)	62.3±8.6	(49–79)
APS e	(Range)	1.2±0.9	(0.6–2.2)	0.2±0.3	(0–0.6)	3.2±2.2	(2.3–9.11)
Ntp f	(Range)	1.6±0.5	(1–2)	0.4±0.8	(0–2)	1.3±0.7	(0–2)
APT g (%)	(Range)	11.9±4.3	(6.7–16.7)	3.0±5.2	(0–12.5)	8.2±5.2	(0–16.7)
Body wall	Anchor (n)	8		8		12
Length	(Range)	165.4±5.6	(160–174)	160.4±5.9	(153–173)	155.4±6.0	(143–164)
Width	(Range)	64.8± 4.5	(57–71)	68.0±3.2	(61–71)	76.9±4.3	(69–84)
Teeth	(Range)	6.5±0.9	(5–8)	4.0±1.3	(1–5)	9.3±1.5	(7–12)
ASW h(%)	(Range)	27.8±2.0	(25.8–32.5)	24.7±2.7	(21.8–29.5)	20.3±0.9	(19.1–21.6)
AEW i (%)	(Range)	55.1±5.2	(50.0–63.2)	51.8±5.0	(45.1–60.7)	48.9±2.0	(45.6–53.1)
Dorsal side
L-Muscle	Granules (n)	12		14		11
Length	(Range)	32.5±4.7	(28–42)	28.7±3.9	(21–34)	27.9±6.3	(21–39)
Width	(Range)	13.5±2.0	(9–17)	13.8±1.7	(11–16)	15.0±1.8	(12–18)
GP c	(Range)	42.4±9.1	(28.2–56.7)	48.6±6.6	(35.3–59.1)	56.7±15.9	(35.3–81.8)
GCO d (%)	(Ratio)	33.3	(O4: C8)	78.6	(O11: C3)	54.5	(O6: C5)
Body wall	Anchor plates (n)	8		9		10
Length	(Range)	93.9±4.2	(88–99)	89.4±8.1	(75–98)	94.5±3.8	(88–99)
Width	(Range)	50.4±3.0	(47–55)	55.6±2.8	(53–62)	59.1±5.7	(49–67)
APS e	(Range)	0.7±0.6	(0–2.1)	0.2±0.3	(0–0.7)	1.8 ±1.6	(0.5–5.4)
Ntp f	(Range)	2.5±0.9	(1–4)	1.2±1.0	(0–3)	1.7±0.7	(1–3)
APT g (%)	(Range)	16.8 ± 4.9	(8.3–23.1)	10.1 ± 8.8	(0–27.3)	9.8±3.3	(5.3–13.6)
Body wall	Anchor (n)	8		12		10
Length	(Range)	162.3 ± 8.5	(145–172)	153.1 ± 7.9	(144–168)	157.4±6.7	(145–168)
Width	(Range)	61.5 ± 4.7	(54–67)	64.8 ± 3.2	(58–70)	79.7±4.9	(73–89)
Teeth	(Range)	6.2 ± 0.6	(5–7)	3.2 ± 1.9	(0–6)	9.9±1.2	(8–12)
ASW h(%)	(Range)	28.4 ± 2.5	(25.8–32.4)	25.0 ± 1.9	(22.2–28.1)	19.9±1.1	(18.5–22.1)
AEW i (%)	(Range)	59.2 ± 4.3	(52.2–64.9)	50.8 ± 4.7	(41.2–55.6)	48.6±2.7	(43.8–52.0)