Rediscovery and revalidation of Calamaria annamensis Bourret, 1937 (Squamata: Calamariidae), a rare reed snake restricted to the Truong Son Range of central Vietnam Author Korolev, Alexey M. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119234, Russia Author Bragin, Andrey M. Joint Vietnam-Russia Tropical Science and Technology Research Center, Nghia Do, Hanoi, Vietnam Author Gorin, Vladislav A. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119234, Russia Author Le, Son Xuan Joint Vietnam-Russia Tropical Science and Technology Research Center, Nghia Do, Hanoi, Vietnam Author Idiiatullina, Sabira S. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119234, Russia Author Xu, Yuhao State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining, China Author Zhang, Tierui Anhui Province Key Laboratory of the Conservation and Exploitation of Biological Resource, College of Life Sciences, Anhui Normal University, Wuhu, China Author Bos, Collin Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720 - 3160, U. S. A. Author David, Patrick Institut de Systématique, Évolution et Biodiversité (ISYEB), Muséum National d’Histoire Naturelle, Sorbonne Université, École Pratique des Hautes Études, Université des Antilles, CNRS, CP 30, 57 rue Cuvier, F- 75005 Paris, France Author Nguyen, Tan Van The School of Medicine & Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam & Center for Entomology & Parasitology Research, Duy Tan University, Da Nang, 550000, Vietnam Author Poyarkov, Nikolay A. Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119234, Russia & Joint Vietnam-Russia Tropical Science and Technology Research Center, Nghia Do, Hanoi, Vietnam text Zootaxa 2026 2026-01-26 5752 1 105 133 https://doi.org/10.11646/zootaxa.5752.1.5 journal article 315344 10.11646/zootaxa.5752.1.5 8dbb956d-ce9b-4057-ae94-d16a3d9e4785 1175-5326 19171592 A8C82F70-EF6F-4586-97AF-F246D619A696

TABLE 4. Comparison of morphological characters of Calamaria annamensis stat. nov. with those of congeners occurring in China and the Indo-Burma region. Symbols: (1) = Supralabials; (2) = Supralabials contacting orbit; (3) = Preocular (1 = present, 0 = absent); (4) = Mental touching chin shields (1 = yes, 0 = no); (5) = Number of scales contacting paraparietal; (6) = VEN in males; (7) = VEN in females; (8) = SC in males; (9) = SC (females); (10) = maximum TL (mm) in males; (11) = maximum TL (mm) in females; (12) = ratio TaL/TL (%) in males; (13) = ratio TaL/TL (%) in females. Notes: N/a: Not available; diagnostic characters (with respect to Calamaria annamensis stat. nov.) are highlighted in bold.

Species	(1)	(2)	(3)	(4)	(5)	(6)	(7)	(8)	(9)	(10)	(11)	(12)	(13)
C. annamensis stat. nov.	4	2/3	1	0	6	182	196–204	22	14–15	304	552	6.6	3.9–4.7
C. abramovi	4	2/3	1	0	6	159	174	26	20	139	482	13.3	7.1
C. andersoni	4	2/3	1	0	6	164–172	186	20–23	n/a	351	312	8.8–9.2	5.8
C. arcana	4	2/3	1	0	6	170–176	192	20–22	12	303	n/a	7.2–11.8	4.7
C. berezowskii	4	2/3	1	0	6	149–155	153–171	22–25	12–16	290	245	6.6–10.5	6.5–6.9
C. buchi	4	2/3	1	1	5	n/a	221–236	n/a	13–14	n/a	466	n/a	3.9–4.1
C. concolor	5	2/3	1	1	5	209	n/a	19	n/a	578	n/a	7.2	n/a
C. dominic	4 or 5	2/3 or 3/4	1	1 or 0	6	n/a	174	n/a	17 or 18	n/a	421	n/a	6.2
C. gialaiensis	4	2/3	1	1	5	191	n/a	23	n/a	457	n/a	8.1	n/a
C. jinggangensis	4	2/3	1	0	6	157–158	179	20	12	260	364	15	3.6
C. lovii ingermarxorum	4	2/3	0	1	6	205	n/a	23	n/a	318	n/a	7.4	n/a
C. lumbricoidea	5	3/4	1	1	4 or 5	144–196	137–229	17–27	13–21	498	642	6.3–11.4	3.9–8.3
C. mizoramensis	4	2/3	1	0	6	147–155	166–175	22–27	13–15	251	258	10.3–13.1	4.7–6.5
C. nebulosa	4	2/3	0	0	6	n/a	179	n/a	22	n/a	354	n/a	7.9
C. pavimentata	4	2/3	1	0	n/a	151	136	27	13	n/a	n/a	n/a	n/a
C. sangi	4	2/3	1	1	5 or 6	190	n/a	19	n/a	373	n/a	6.2	n/a
C. schlegeli	5	3/4	1 or 0	0	5 or 6	129–161	136–180	25–44	19–37	391	395	11.1–21.3	7.3–14.4
C. septentrionalis	4	2/3	1	0	6	148–166	168–188	15–19	6–11	344	384	6.3–8.6	2.6–4.3
C. strigiventris	4	2/3	1	0	6	130–168	176–183	29–31	20–22	362	367	11.2–17.9	8.4–8.6
C. synergis	4	2/3	1	0	5	161–166	n/a	20–23	n/a	284	n/a	6.6–9.2	n/a
C. thanhi	4	2/3	0	0	6 or 7	184	198	28	21	461	455	9.9	6.8
C. yunnanensis	4	2/3	0	0	6	167–184	199	15–20	19	300	516	8.2–8.8	5.4–8.2