A new species of limestone-forest frog, genus Platymantis (Amphibia: Anura: Ceratobatrachidae) from central Luzon Island, Philippines Author Siler, Cameron D. Author Diesmos, Arvin C. Author Linkem, Charles W. Author Diesmos, Mae L. Author Brown, Rafe M. text Zootaxa 2010 2482 49 63 journal article 48154 10.5281/zenodo.195468 e2290a51-8f8f-4a97-95a9-5a8710c4ece4 1175-5326 195468

TABLE 1. Summary of qualitative diagnostic characters (present, absent) in Platymantis biak and specimens of other members of the P. guentheri Species Group. Sample size for each sex, body size, eye diameter, digit disk proportions, and general geographical distribution (PAIC = Pleistocene Aggregate Island Complexes, sensu Brown and Diesmos, 2002) are included for reference (SVL and ED given as range over mean ± standard deviation; digit disk proportions given as percentage over mean ± standard deviation). Only adult specimens in good preservation condition were used for morphometric analyses.

	biak sp. nov. (24m, 8f)	bayani (9m, 9f)	insulatus (8m, 6f)	negrosensis (3m, 2f)	luzonensis (9m, 2f)	rabori (2m, 6f)	guentheri (11m, 8f)	banahao (4m)	cornutus (3m)	diesmosi (8m, 2f)
Range	Central Luzon Island	Samar Island	Gigantes Islands	Negros and Panay Islands	Luzon PAIC	Mindanao PAIC	Mindanao PAIC	SE Luzon Island	N. Luzon Island	SE Luzon Island
SVL (m) SVL (f)	32.3–39.9 (35.9 ± 1.8) 37.4–42.4 (40.1 ± 1.5)	34.2–39.1 (37.1 ± 1.8) 44.3–49.8 (46.4 ± 2.1)	34.3–38.0 (36.7 ± 1.3) 40.2–42.9 (41.8 ± 1.0)	28.3–35.7 (31.1 ± 4.0) 41.7–42.2 (42.0 ± 0.3)	24.1–32.5 (29.7 ± 2.4) 32.6–33.8 (33.2 ± 0.8)	30.4–33.5 (32.0 ± 2.2) 36.4–44.4 (41.5 ± 3.6)	26.5–29.8 (28.6 ± 1.1) 32.3–37.3 (34.6 ± 2.0)	32.4–36.3 (33.9 ± 1.7) N/A	26.3–30.4 (27.8 ± 2.2) N/A	28.5–39.2 (35.3 ± 3.6) 49.0–52.6 (50.8 ± 2.6)
ED (m) ED (f)	3.3–5.2 (4.3 ± 0.5) 4.3–5.2 (4.9 ± 0.3)	4.8–6.1 (5.5 ± 0.5) 5.4–6.8 (6.1 ± 0.4)	3.7–4.9 (4.2 ± 0.4) 5.1–5.7 (5.3 ± 0.2)	3.1–4.1 (3.7 ± 0.6) 4.4–4.7 (4.6 ± 0.2)	2.7–3.9 (3.4 ± 0.4) 3.6–3.9 (3.7 ± 0.2)	3.8–3.9 (3.9 ± 0.1) 4.0–5.7 (4.9 ± 0.7)	2.8–3.6 (3.3 ± 0.3) 3.7–4.5 (3.9 ± 0.3)	3.6–4.1 (3.8 ± 0.2) N/A	3.0–4.2 (3.5 ± 0.6) N/A	3.7–5.4 (4.7 ± 0.5) 5.9–6.1 (6.0 ± 0.1)
Fin3DW/PpFin3 (m) Fin3DW/PpFin3 (f)	1.4–3.4 (2.1 ± 0.5) 1.4–2.7 (2.2 ± 0.4)	2.7–4.0 (3.3 ± 0.4) 2.8–4.5 (3.9 ± 0.5)	2.7–3.1 (2.9 ± 0.1) 2.3–2.8 (2.7 ± 0.2)	2.6–2.8 (2.7 ± 0.1) 3.5–4.7 (4.1 ± 0.8)	2.2–3.3 (2.8 ± 0.4) 3.0–3.2 (3.1 ± 0.2)	2.8–3.5 (3.2 ± 0.5) 3.3–4.2 (3.6 ± 0.3)	2.4–5.2 (3.2 ± 0.8) 2.6–5.7 (4.0 ± 1.2)	2.8–3.2 (3.0 ± 0.2) N/A	2.5–2.9 (2.7 ± 0.2) N/A	1.5–2.1 (1.7 ± 0.2) 1.6–1.7 (1.6 ± 0.1)
Toe4DW/PpToe4 (m) Toe4DW/PpToe4 (f)	1.3–2.4 (1.7 ± 0.3) 1.6–3.1 (2.1 ± 0.5)	2.1–3.3 (2.8 ± 0.4) 2.9–3.7 (3.1 ± 0.3)	1.9–3.4 (2.6 ± 0.4) 2.0–2.6 (2.3 ± 0.3)	1.8–1.9 (1.8 ± 0.1) 1.8–1.8 (1.8 ± 0.0)	1.6–2.2 (1.9 ± 0.2) 1.8–1.9 (1.8 ± 0.1)	2.0–2.7 (2.3 ± 0.5) 1.6–2.5 (2.3 ± 0.3)	1.5–3.5 (2.4 ± 0.6) 2.3–4.3 (3.2 ± 0.7)	1.9–2.3 (2.0 ± 0.2) N/A	1.6–2.0 (1.7 ± 0.2) N/A	1.2–2.2 (1.7 ± 0.4) 1.7–2.1 (1.9 ± 0.3)
Fin3DW/ Toe4DW (m) Fin3DW/Toe4DW (f)	0.9–1.7 (1.2 ± 0.2) 0.9–1.4 (1.2 ± 0.2)	1.1–1.5 (1.3 ± 0.1) 1.2–1.5 (1.3 ± 0.1)	1.1–1.5 (1.3 ± 0.1) 1.2–1.6 (1.4 ± 0.1)	1.8–2.0 (1.9 ± 0.1) 1.7–2.0 (1.9 ± 0.3)	1.6–2.0 (1.7 ± 0.1) 2.0–2.1 (2.1 ± 0.1)	1.3–1.6 (1.4 ± 0.2) 1.4–1.9 (1.6 ± 0.2)	1.2–1.7 (1.4 ± 0.1) 1.1–1.7 (1.4 ± 0.2)	1.8–1.9 (1.8 ± 0.0) N/A	1.9–2.1 (2.0 ± 0.1) N/A	0.9–1.4 (1.1 ± 0.2) 0.8–0.8 (0.8 ± 0.0)
Greatly expanded digital disks	+, moderate	+	+	+	+	+	+, moderate	+	+	–
Dorsal skin rugosity	–	– to granular	–	+	+	+	+	– to granular	–	– to granular
Dorsal dermal tubercles	–	+, sparse	–	+	+	+, dense	+	+	–	–
Tubercle color	N/A	Salmon	N/A	Cream	Dominant dorsal color	Dominant dorsal color	Dominant dorsal color	Dominant dorsal color	N/A	N/A
Dorsal folds/ridges	–	–	–	+	+	+	+	–	–	–
Tympanum hidden	No	No	No	Posterodorsal edge	Posteroventral edge	Posterior edge	Posteroventral edge	Posterodorsal edge	No	Posterodorsal edge
Supratympanic fold	Smooth, less prominent	Smooth, less prominent	Smooth, not prominent	Rugose, prominent	Rugose, prominent	Rugose, prominent	Smooth, less prominent	Smooth, less prominent	Smooth, not prominent	Smooth, less prominent
Dorsolateral stripes	–	–	–	+	+ or –	+ or –	+ or –	–	+	–
Spotted flanks	–, blotches anteriorly	+	+	–	–	–	–	–	–	–
Limb bands	+, well defined	+, well defined	+ or –, poorly defined	+, well defined	+, well defined	+, well defined	+, well defined	+ or –, faint	+, poorly defined	+, well defined
Microhabitat preference	Ground, karst habitat	Ground; karst habitat	Ground; karst habitat	Arboreal	Arboreal	Arboreal	Shrub/Arboreal	Arboreal	Arboreal	Ground; cliff edges