Benstonea Callm. & Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species Author Callmander, Martin W. Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Conservatoire et Jardin botaniques de la Ville de Genève, CP 60, CH- 1292 Chambésy, Switzerland. martin.callmander@mobot.org Author Lowry Ii, Porter P. Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Département Systématique et Evolution (UMR 7205), Muséum National d’Histoire Naturelle, case postale 39, rue Cuvier 57, 75231 Paris (Cedex) 05, France. Author Forest, Félix JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom. Author Devey, Dion S. JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom. Author Beentje, Henk Herbarium, Library, Art & Archives, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom. Author Buerki, Sven JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom. text Candollea 2012 2012-12-01 67 2 323 345 journal article 10.15553/c2012v672a12 2235-3658 5790002

Appendix 1. – Complete list of accepted names in Benstonea Callm. & Buerki with synonyms (incl. basionyms).

	Accepted taxon name	Synonyms (incl. Basionyms)
1	Benstonea adinobotrys (Merr. & L. M. Perry) Callm. & Buerki	P. adinobotrys Merr. & L. M. Perry P. angiensis Kaneh. P. congregatus H. St. John P. ketele B. C. Stone P. lustrorum B. C. Stone
2	Benstonea affinis (Kurz) Callm. & Buerki	P. affinis Kurz P. aurantiacus Ridl. P. merrillii Warb. P. sarawakensis Martelli
3	Benstonea alticola (Holttum & H. St. John) Callm. & Buerki	P. alticola Holttum & H. St. John
4	Benstonea andersonii (H. St. John) Callm. & Buerki	P. andersonii H. St. John
5	Benstonea ashtonii (B. C. Stone) Callm. & Buerki	P. ashtonii B. C. Stone
6	Benstonea atrocarpa (Griff.) Callm. & Buerki	P. atrocarpus Griff.
7	Benstonea beccata (B. C. Stone) Callm. & Buerki	P. beccatus B. C. Stone
8	Benstonea biplicata (H. St. John) Callm. & Buerki	P. biplicatus H. St. John
9	Benstonea brevistylis (B. C. Stone) Callm. & Buerki	P. brevistylis B. C. Stone
10	Benstonea brunigii (B. C. Stone) Callm. & Buerki	P. brunigii B. C. Stone
11	Benstonea calcinacta (B. C. Stone) Callm. & Buerki	P. calcinactus B. C. Stone
12	Benstonea celebica (Warb.) Callm. & Buerki	P. celebicus Warb.
13	Benstonea copelandii (Merr.) Callm. & Buerki	P. copelandii Merr. P. copelandii var. panchoi B. C Stone P. muricatus Merr.
14	Benstonea ellipsoidea (Warb.) Callm. & Buerki	P. ellipsoideus Warb. P. dipsaceus Martelli P. kjellbergii Fagerl.
15	Benstonea elostigma (Martelli) Callm. & Buerki	P. elostigma Martelli
16	Benstonea foetida (Roxb.) Callm. & Buerki	P. foetidus Roxb. P. wallichianus Martelli, inval. name
17	Benstonea gibbsiana (Martelli) Callm. & Buerki	P. gibbsianus Martelli
		P. obovoideus Merr.
18	Benstonea glaucophylla (Ridl.) Callm. & Buerki	P. glaucophyllus Ridl.
19	Benstonea herbacea (Martelli) Callm. & Buerki	P. herbaceus Martelli
		P. collinus Ridl.
		P. collinus var. spinulosus Ridl.
		P. kingianus Martelli
20	Benstonea humilis (Lour.) Callm. & Buerki	P. humilis Lour.
		Fisquetia ovata Gaudich., inval. name
		P. distans Craib
		P. leucocephalus Martelli
		P. ovatus Kurz, inval. name
		P. ovatus Warb.
		P. perakensis Ridl.
		P. pierrei Martelli
		P. pierrei var. bariensis Martelli
		P. pierrei var. constrictus Gagn.
		P. pseudofoetidus Martelli
		P. retroaculeatus H. St. John
		P. similis Craib
		P. toinu H. St. John
21	Benstonea inquilina (B. C. Stone) Callm. & Buerki	P. inquilinus B. C. Stone
22	Benstonea korthalsii (Solms) Callm. & Buerki	P. korthalsii Solms
23	Benstonea kurzii (Merr.) Callm. & Buerki	P. kurzii Merr.
		P. asper H. St. John
24	Benstonea lauterbachii (K. Schum. & Warb.) Callm. & Buerki	P. lauterbachii K. Schum. & Warb.
25	Benstonea lepatophila (B. C. Stone) Callm. & Buerki	P. lepatophilus B. C. Stone
26	Benstonea monticola (F. Muell.) Callm. & Buerki	P. monticola F. Muell. P. pluvisilvaticus H. St. John
27	Benstonea nana (Martelli) Callm. & Buerki	P. nanus Martelli
		P. magnifibrosus H. St. John
28	Benstonea odoardii (Martelli) Callm. & Buerki	P. odoardii Martelli P. eumekes B. C. Stone P. humicola Kaneh. P. ihuanus Martelli P. inokumae Kaneh. P. papuanus Ridl., illeg. name P. pentagonos H. St. John P. rostellatus Merr. & L. M. Perry
29	Benstonea ornata (Solms) Callm. & Buerki	P. ornatus Solms Fisquetia ornata Gaudich., inval. name P. monotheca Martelli P. ornatus Kurz, inval. name P. ornatus f. spicatus Martelli P. recurvatus H. St. John
30	Benstonea pachyphylla (Merr.) Callm. & Buerki	P. pachyphyllus Merr. P. apicalis H. St. John
31	Benstonea parva (Ridl.) Callm. & Buerki	P. parvus Ridl. P. flagellifer Warb. P. mollifoliaceus H. St. John
32	Benstonea pectinata (Martelli) Callm. & Buerki	P. pectinatus Martelli P. papilio B. C. Stone
33	Benstonea permicron (Kaneh.) Callm. & Buerki	P. permicron Kaneh.
34	Benstonea pilaris (Ridl.) Callm. & Buerki	P. pilaris Ridl.
35	Benstonea poronavila (B. C. Stone) Callm. & Buerki	P. poronavila B. C. Stone
36	Benstonea pseudosyncarpa (Kaneh.) Callm. & Buerki	P. pseudosyncarpus Kaneh.
37	Benstonea pumila (H. St. John) Callm. & Buerki	P. pumilus H. St. John
38	Benstonea rupestris (B. C. Stone) Callm. & Buerki	P. rupestris B. C. Stone
		P. scandens B. C. Stone
		P. oresbios B. C. Stone
39	Benstonea rustica (B. C. Stone) Callm. & Buerki	P. rusticus B. C. Stone
40	Benstonea setistyla (Warb.) Callm. & Buerki	P. setistylus Warb.
41	Benstonea stenocarpa (Solms) Callm. & Buerki	P. stenocarpus Solms
		P. acicularis H. St. John
		P. arcuatus H. St. John
		P. danckelmannianus K. Schum.
		P. erinaceus B. C. Stone
		P. gladiator B. C. Stone
		P. jacobsii B. C. Stone
		P. lictor B. C. Stone
		P. misimaensis B. C. Stone
		P. nigridens B. C. Stone
		P. verruculosus B. C. Stone
42	Benstonea sylvatica (B. C. Stone) Callm. & Buerki	P. sylvaticus B. C. Stone
43	Benstonea thomissophylla (B. C. Stone) Callm. & Buerki	P. thomissophyllus B. C. Stone
44	Benstonea thurstonii (C. H. Wright) Callm. & Buerki	P. thurstonii C. H. Wright P. varawa H. St. John
45	Benstonea thwaitesii (Martelli) Callm. & Buerki	P. thwaitesii Martelli P. foetidus var. racemosus Trimen
46	Benstonea toei (H. St. John) Callm. & Buerki	P. toei H. St. John
47	Benstonea tunicata (B. C. Stone) Callm. & Buerki	P. tunicatus B. C. Stone
48	Benstonea undulifolia (Holttum & H. St. John) Callm. & Buerki	P. undulifolius Holttum & H. St. John
49	Benstonea unguiculata (Ridl.) Callm. & Buerki	P. unguiculatus Ridl. P. attenuatus H. St. John P. caudatifolius H. St. John P. globuliferus Ridl. P. globulosus H. St. John
50	Benstonea vriensii (Martelli) Callm. & Buerki	P. vriensii Martelli