Cryptic diversity in toads of the Rhinella marina species group (Anura, Bufonidae) with a subjectively beautiful new species from Western Ecuador Author Menéndez-Guerrero, Pablo A. Laboratorio de Macroecología y Cambio Global, Facultad de Ciencias Exactas y Naturales, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado Postal 17 - 01 - 2184, Quito, Ecuador & Redpath Museum, McGill University, 859 Sherbrooke Street West, Montreal, QC, H 3 A 2 K 6, Canada Author Lima, Sueny P. Author Santos Museo de Zoología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado Postal 17 - 01 - 2184, Quito, Ecuador & Translational Biomedical Sciences, Ohio University, 02 Grosvenor Hall, Athens, OH, USA Author Salazar-Nicholls, María-José Museo de Zoología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado Postal 17 - 01 - 2184, Quito, Ecuador & Department of Biology, Boston University, 5 Cummington Mall, Boston, MA 02215, USA Author Green, David M. Redpath Museum, McGill University, 859 Sherbrooke Street West, Montreal, QC, H 3 A 2 K 6, Canada Author Ron, Santiago R. Museo de Zoología, Pontificia Universidad Católica del Ecuador, Avenida 12 de Octubre 1076 y Roca, Apartado Postal 17 - 01 - 2184, Quito, Ecuador text Zoological Journal of the Linnean Society 2024 2024-02-20 202 3 1 26 https://doi.org/10.1093/zoolinnean/zlad197 journal article 306574 10.1093/zoolinnean/zlad197 046a1646-6c0e-4090-b2fb-1c9a49970aa5 0024-4082 14423165 7-01-2184

Table 3. Descriptive statistics for call parameters of toad populations form Central America, western Ecuador, and eastern Ecuador. Sample size (N), and mean ± SD are given. Individuals from western Ecuador correspond to R. bella, individuals from eastern Ecuador correspond to R. marina, and individuals from Central America correspond to R. horribilis

Variables	Central America	Western Ecuador	Eastern Ecuador
	(N = 5)	(N = 17)	(N = 12)
Call length	6.33 ± 1.01	1.51 ± 0.43	5.28 ± 4.04
Call rise time	3.75 ± 1.86	0.94 ± 0.26	3.09 ± 2.19
Notes per call	83.98 ± 11.10	18.33 ± 5.70	81.66 ± 61.97
Pulses per note	3.55 ± 0.78	3.80 ± 0.59	3.16 ± 0.69
Pulse rate	90.57 ± 13.87	91.31 ± 11.70	88.92 ± 13.66
Call repetition rate	4.41 ± 1.61	8.57 ± 2.40	2.96 ± 0.86
Dominant frequency	633.02 ± 35.28	803.05 ± 75.7	705.52 ± 117.24