Revision of zigzag geckos (Diplodactylidae: Amalosia) in eastern Australia, with description of five new species Author Hoskin, Ad. J. Author Couper, Patrick J. Biodiversity & Geosciences Program, Queensland Museum, South Brisbane, QLD 4101, Australia text Zootaxa 2023 2023-09-11 5343 4 301 337 http://dx.doi.org/10.11646/zootaxa.5343.4.1 journal article 267061 10.11646/zootaxa.5343.4.1 d4e33531-7bd1-4669-8dff-561337c3dd6b 1175-5326 8334387 358CE9AF-F36D-4EEA-89E6-8B64FEFE0772

TABLE 1. Measurements, proportions, and scale counts for Amalosia species of southern Queensland and New South Wales; taken from specimens listed in the type series and material examined for each species. Data are shown as mean followed by range in brackets. Qualitative scale traits are summarized as the typical state or states. Asterisks denote higher N for some original tail measures (see Methods). Male pore and pore gap counts were supplemented with field data (from macro photos) for A. hinesi sp. nov., A. jacovae and A. lesueurii.

	A. queenslandia sp. nov.	A. hinesi sp. nov.	A. jacovae	A. lesueurii	A. cf. rhombifer	A. capensis sp. nov.	A. queenslandia sp. nov.	A. nebula sp. nov.	A. saxicola sp. nov.
TABLE 1. Measurements, proportions, and scale counts for Amalosia species of southern Queensland and New South Wales; taken from specimens listed in the type series and material examined for each species. Data are shown as mean followed by range in brackets. Qualitative scale traits are summarized as the typical state or states. Asterisks denote higher N for some original tail measures (see Methods). Male pore and pore gap counts were supplemented with field data (from macro photos) for A. hinesi sp. nov., A. jacovae and A. lesueurii.
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Measurements	N = 27	N = 13	N = 10	N = 10	N = 11	N = 15	N =27	N =16	N = 23
SVL	44.6 (38.9–50.4)	55.5 (48.5–59.9)	54.2 (49.6–59.5)	54.0 (46.7–58.5)	47.0 (41.6–52.8)	45.0 (40.2–50.7)	44.6 (38.9–50.4)	52.3 (46.4–58.0)	58.2 (52.4–64.6)
Interlimb (AG)	21.5 (18.6–25.2)	25.2 (21.1–30.6)	24.2 (21.0–26.9)	24.3 (21.6–28.1)	22.6 (18.4–26.1)	21.3 (18.3–25.5)	21.5 (18.6–25.2)	23.8 (20.2–28.0)	25.5 (21.0–29.5)
Forelimb (L1)	12.3 (10.0–14.8)	15.6 (14.0–17.9)	15.5 (12.3–17.5)	15.3 (14.1–16.5)	13.5 (11.8–15.9)	11.8 (9.9–13.2)	12.3 (10.0–14.8)	14.9 (13.0–17.2)	16.2 (14.0–19.1)
Hindlimb (L2)	15.5 (13.5–17.4)	19.6 (17.7–21.8)	19.2 (18.2–21.2)	19.4 (18.1–21.1)	16.6 (14.5–20.1)	15.4 (13.2–18.8)	15.5 (13.5–17.4)	18.5 (15.8–20.6)	21.8 (18.1–25.1)
Body width (BW)	7.2 (5.0–8.7)	10.3 (8.6–12.9)	9.8 (8.1–11.3)	11.0 (9.2–12.5)	8.0 (5.6–9.6)	7.4 (6.1–9.9)	7.2 (5.0–8.7)	9.0 (7.0–12.2)	10.7 (9.3–14.6)
Head length (HL)	9.8 (8.5–11.2)	12.3 (11.4–13.4)	11.9 (11.3–12.9)	12.5 (11.5–13.3)	10.3 (9.2–11.3)	9.7 (7.5–10.8)	9.8 (8.5–11.2)	11.4 (9.7–12.6)	13.3 (11.7–15.7)
Head width (HW)	7.3 (6.3–8.5)	9.8 (8.9–10.5)	9.0 (8.2–10.0)	10.2 (9.4–10.8)	7.6 (6.9 –8.5)	7.3 (6.6–7.9)	7.3 (6.3–8.5)	8.7 (7.7–9.8)	10.2 (9.3–11.7)
Head depth (HD)	3.9 (3.1–4.8)	5.2 (4.3–6.1)	4.8 (4.2–5.2)	5.3 (4.5–5.8)	4.6 (3.8–5.4)	3.9 (3.1–4.5)	3.9 (3.1–4.8)	4.6 (3.9–5.6)	4.7 (4.0–6.1)
Snout (S)	4.0 (3.3–4.8)	5.2 (4.5–5.7)	5.2 (4.7–5.8)	5.4 (4.9–5.9)	4.2 (3.5–4.9)	4.2 (3.6–4.8)	4.0 (3.3–4.8)	4.9 (4.4–5.3)	5.9 (5.1–7.0)
Proportions	N = 27	N = 13	N = 10	N = 10	N = 11	N = 15	N = 27	N = 16	N = 23
AG/SVL	0.48 (0.44–0.55)	0.45 (0.41–0.51)	0.45 (0.40–0.48)	0.45 (0.41–0.48)	0.48 (0.43–0.52)	0.47 (0.42–0.52)	0.48 (0.44–0.55)	0.46 (0.40–0.49)	0.44 (0.39–0.48)
L1/SVL	0.27 (0.24–0.31)	0.28 (0.25–0.30)	0.29 (0.25–0.32)	0.28 (0.25–0.33)	0.29 (0.24–0.33)	0.26 (0.22–0.29)	0.27 (0.24–0.31)	0.28 (0.26–0.30)	0.28 (0.26–0.31)
L2/SVL	0.35 (0.28–0.38)	0.35 (0.33–0.41)	0.35 (0.33–0.39)	0.36 (0.34–0.39)	0.36 (0.29–0.38)	0.34 (0.29–0.39)	0.35 (0.28–0.38)	0.35 (0.33–0.38)	0.38 (0.33–0.42)
BW/SVL	0.16 (0.12–0.19)	0.18 (0.16–0.22)	0.18 (0.16–0.21)	0.20 (0.18–0.23)	0.17 (0.14–0.20)	0.17 (0.14–0.22)	0.16 (0.12–0.19)	0.17 (0.15–0.21)	0.18 (0.15–0.23)
HL/SVL	0.22 (0.20–0.23)	0.22 (0.21–0.23)	0.22 (0.21–0.23)	0.23 (0.22–0.25)	0.22 (0.21–0.23)	0.22 (0.19–0.23)	0.22 (0.20–0.23)	0.22 (0.20–0.23)	0.23 (0.21–0.25)
HW/SVL	0.16 (0.15–0.18)	0.18 (0.17–0.18)	0.17 (0.16–0.18)	0.19 (0.18–0.20)	0.16 (0.15–0.17)	0.16 (0.15–0.18)	0.16 (0.15–0.18)	0.17 (0.16–0.17)	0.18 (0.16–0.19)
HD/SVL	0.09 (0.07–0.11)	0.09 (0.07–0.12)	0.09 (0.07–0.10)	0.10 (0.09–0.12)	0.10 (0.09–0.11)	0.09 (0.07–0.10)	0.09 (0.07–0.11)	0.09 (0.07–0.10)	0.08 (0.07–0.11)
HW/HL	0.75 (0.68–0.88)	0.79 (0.74–0.84)	0.76 (0.71–0.83)	0.81 (0.78–0.86)	0.73 (0.69–0.78)	0.75 (0.69–0.90)	0.75 (0.68–0.88)	0.77 (0.71–0.82)	0.77 (0.70–0.87)
HD/HL	0.40 (0.32–0.49)	0.42 (0.34–0.51)	0.40 (0.35–0.45)	0.43 (0.37–0.50)	0.45 (0.39–0.49)	0.40 (0.34–0.51)	0.40 (0.32–0.49)	0.40 (0.31–0.48)	0.36 (0.30–0.50)
HD/HW	0.54 (0.44–0.66)	0.53 (0.44–0.66)	0.53 (0.43–0.57)	0.52 (0.46–0.62)	0.61 (0.54–0.69)	0.54 (0.46–0.62)	0.54 (0.44–0.66)	0.53 (0.40–0.61)	0.47 (0.40–0.61)
S/HL	0.40 (0.36–0.46)	0.42 (0.39–0.46)	0.43 (0.41–0.49)	0.43 (0.40–0.45)	0.41 (0.37–0.43)	0.43 (0.40–0.47)	0.40 (0.36–0.46)	0.43 (0.41–0.46)	0.44 (0.40–0.47)
Original Tail	N = 6 (18*)	N = 3 (7*)	N = 2 (5*)	N = 4 (6*)	N = 4 (7*)	N = 3 (10*)	N = 6 (18*)	N = 5 (12*)	N = 12 (21*)
Tail Length (oTL)	42.4 (36.4–47.0)	50.8 (46.0–53.5)	55.9 (55.8–56.0)	43.0 (40.3–45.7)	46.8 (43.2–51.6)	38.2 (35.1–41.1)	42.4 (36.4–47.0)	48.0 (44.4–53.2)	49.7 (43.9–54.0)
Tail Width (oTW)*	4.2 (3.3–5.3)	5.7 (5.2–6.3)	6.0 (5.0–7.4)	7.3 (6.1–8.1)	4.3 (3.5–5.2)	4.2 (3.2–4.7)	4.2 (3.3–5.3)	5.0 (3.9–5.8)	5.5 (4.3–7.12)
Tail Depth (oTD)*	3.5 (2.7–4.5)	4.8 (4.2–5.4)	4.5 (3.7–5.9)	5.0 (4.5–5.4)	4.0 (3.4–4.3)	3.6 (2.7–4.0)	3.5 (2.7–4.5)	4.2 (3.3–5.4)	4.5 (3.8–5.1)
oTL/SVL	0.95 (0.90–1.01)	0.92 (0.89–0.94)	0.98 (0.94–1.02)	0.83 (0.78–0.91)	0.98 (0.94–1.00)	0.89 (0.87–0.91)	0.95 (0.90–1.01)	0.95 (0.83–1.05)	0.85 (0.80–0.92)
oTW/oTL	0.09 (0.07–0.11)	0.11 (0.10–0.11)	0.11 (0.10–0.13)	0.17 (0.15–0.18)	0.09 (0.07–0.11)	0.10 (0.09–0.11)	0.09 (0.07–0.11)	0.10 (0.09–0.11)	0.11 (0.09–0.14)
oTW/oTD*	1.19 (1.04–1.44)	1.18 (1.07–1.30)	1.35 (1.17–1.76)	1.46 (1.34–1.57)	1.09 (0.96–1.20)	1.19 (1.04–1.55)	1.19 (1.04–1.44)	1.20 (1.07–1.63)	1.23 (1.03–1.43)
Scales	N = 40	N = 14	N = 11	N = 8	N = 11	N = 20	N = 40	N = 17	N = 27
Spurs (one side)	3.2 (2–5)	2.9 (2–4)	3.5 (3–5)	4.9 (4–6)	1.7 (1–2)	3.0 (2–4)	3.2 (2–5)	3.0 (2–4)	4.6 (3–6)
Rostral crease %	0.63 (0.40–0.80)	0.66 (0.50–0.80)	0.49 (0.30–0.60)	0.50 (0.40–0.60)	0.55 (0.40–0.70)	0.56 (0.35–0.65)	0.63 (0.40–0.80)	0.53 (0.50–0.70)	0.44 (0.25–0.60)
Rostral contacts	2.7 (2–4)	2.9 (2–4)	2.6 (2–3)	2.9 (2–4)	2.9 (2–4)	2.6 (2–3)	2.7 (2–4)	2.8 (2–5)	3.0 (2–4)
Mental contacts	5.2 (3–7)	3.9 (3–5)	4.5 (3–5)	5.0 (4–6)	5.0 (3–6)	3.8 (3–5)	5.2 (3–7)	4.9 (3–7)	4.7 (3–6)
Nasal contacts	5.1 (5–6)	5.9 (5–6)	5.9 (5–6)	6.6 (6–7)	5.2 (5–6)	5.6 (5–6)	5.1 (5–6)	5.4 (5–6)	6.0 (5–7)
Supralabials	9.8 (9–11)	11.0 (10–13)	10.4 (9–12)	10.8 (10–12)	10.2 (10–11)	9.7 (8–11)	9.8 (9–11)	9.9 (8–12)	11.4 (10–14)
Infralabials	10.0 (9–11)	10.9 (10–12)	9.8 (9–12)	9.6 (9–10)	10.2 (10–11)	9.4 (7–11)	10.0 (9–11)	10.3 (9–12)	10.9 (9–14)
Supralab. tallest	1, or 1/2 equal	2, or 1/2 equal	2, or 1/2 equal	1, or 1/2 equal	1	1 or 2, or 1 & 2 equal	1, or 1 & 2 equal	1, or 1 & 2 equal	2, or 1 & 2 equal
Supralab. widest	1, or 1/2 equal	2	2	2	1, or 1 & 2 equal	2	1, or 1 & 2 equal	1 or 2, or 1 & 2 equal	2
4th toe lamellae	6.5 (5–8)	6.9 (6–9)	7.8 (7–9)	7.3 (7–8)	7.3 (6–8)	6.7 (5–8)	6.5 (5–8)	7.1 (6–9)	6.4 (5–8)
4th finger lamellae	6.1 (5–8)	6.4 (5–8)	7.3 (6–8)	7.0 (6–8)	7.0 (5–8)	6.0 (5–7)	6.1 (5–8)	6.4 (5–8)	6.5 (5–9)
3rd/4th toe web	None or minimal	Moderate	Pronounced	Mod.–pronounced	Minimal–moderate	None or minimal	None or minimal	None or minimal	Minimal–moderate
Male pores	N = 28	N = 6	N = 12	N = 12	N = 7	N = 15	N = 28	N = 7	N = 15
Pore count	8.5 (4–14)	6.7 (4–9)	16.4 (13–22)	9.0 (7–13)	16.9 (10–24)	17.1 (13–25)	8.5 (4–14)	3.6 (2–7)	3.9 (0–7)
Pore gap	1.0 (0–2)	4.3 (3–6)	5.1 (2–6)	0.6 (0–2)	3.6 (3–4)	1.9 (1–4)	1.0 (0–2)	3.7 (1–5)	3.9 (0–6)