Kapsulotaenia chisholmae n. sp. (Cestoda: Proteocephalidae), from Varanus spenceri (Reptilia: Varanidae) in Australia Author Hugh I. Jones Discipline of Microbiology, M 502, University of Western Australia, Nedlands, WA 6009, Australia alain.dechambrier@ville-ge.ch Author Alain de Chambrier Muséum d’histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland text Revue suisse de Zoologie 2016 2016-02-03 123 2 209 217 journal article 25079 10.5281/zenodo.155139 5f25bfc0-9c8c-4e03-922d-d4d454257b19 0035-418 155139

Table 1. Summary of diagnostic traits in Kapsulotaenia species. Measurements in micrometres (μm) unless otherwise stated. PC = proportion of the length of cirrus-sac to the width of the proglottis; PP = position of the genital pore (cirrus pore) as % of the proglottis length; OV = percentage of the width of the ovary in relation to the width of the proglottis; SL = supplementary layer within the embryophore (see de Chambrier, 2006, p. 91);* see de Chambrier et al. (2012); D = taken from the original drawings; B = measurements taken from cotypes of Beddard (BMNH 1915.5.14.2); R = measurements taken from type specimens of Ratz (MHNG-PLAT-36037).

Specie s	K. chisholmae n. sp.	K. frezei Schmidt & Kuntz, 1974	K. sangroundi (Carter, 1943)	K. tidswelli (Johnston, 1909)	K. varia (Beddard, 1913)	K. varia (Beddard, 1913) sensu Nybelin (1917) **	K. saccifera (Ratz, 1900) sp. inq.
Reference	This paper	Schmidt & Kuntz, 1974	de Chambrier, 2006	Johnston, 1909; Freze, 1965	Beddard, 1913	Nybelin, 1917; Freze, 1965	Ratz, 1900a, b; Freze, 1965
Host	V. spenceri Lucas & Frost, 1903	V. salvator (Laurenti, 1768) = V. palawanensis Koch, Gaulke & Böhme, 2010	Varanus komodoensis Ouwens 1912	V. varius (White, 1790)	V. varius (White, 1790)	V. gouldii (Gray, 1838)	Varanus sp.
Type locality	Australia, Nelia, Qld	Philippines, Palawan	Indonesia	Australia, Bathurst, NSW	“Zoological Garden”	Mt Tamborine, Queensland	New Guinea
Total length (mm)	180-315	40	92-300	27 to 30	more than 30	230, possibly 300	10-40
Proglottis width	750-840	700	1400-2000	200-350	? (B = 835)	1500-2000	750
Scolex width	340-450	360-465	640-900	320-400	?	400-420	470
Sucker width	90-160	145-165	180-250	80	?	200-220	150
Apical organ width	70-90	40-50	200-250	?	?	130	?
Testis number	89-132	54-60	92-147	about 90	? (B = 104)	96-150 (x=120)	?
Testis diameter	24x40	50-90	20-45	13-22	?	74-110	36-42
Testis field	2	2	2 converging anteriorly	2	2 converging anteriorly	2	?
PC	20-27%	26% (D)	17-27%	21% (D)	17-20% (B = 23%)	17-20% (D)	?
PP	66-77%	43% (D)	61-73%	about 75%(D=59-70%)	about 50% (B = 69%)	about 50%	about 50%
OV	57-65%	61% (D)	42-59%	54%(D)	? (B = 59%)	60% (D)	?
Ovary surface ratio*	6.9-7.5%	7.2%	3.4-4.2%	4.6% (D)	? (B = 5.1%)	4.0%	?
Vagina	mainly anterior (93%)	mainly anterior	posterior	mainly posterior	posterior	anterior/posterior	anterior/posterior
Vaginal sphincter	absent	absent	present	?	absent	present	?
Uterine branches	not visible	absent	21-31 on each side	not visible	?	?	?
Embryophore diameter	37-45	34-36	25-30	19-32	?	?	?
Oncosphere diameter	12-15	?	10-11	?	?	?	?
No. eggs in cluster	8-13	90-100	7-13	?	12-20	?	(R = much more than 100)
Cluster shape	rounded	banana-shaped	rounded	?	rounded	?	(R = banana-shaped)
SL	Yes	Yes	Yes	?	Yes	?	(R=Yes)
Type of uterine formation	Type 1	Type 1	Type 1	Type 1	Type 1	Type 1	Type 1