Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae) Author Espeland, Marianne Author Nakahara, Shinichi Author Zacca, Thamara Author Barbosa, Eduardo P. Author Huertas, Blanca Author Marín, Mario A. Author Lamas, Gerardo Author Benmesbah, Mohamed Author Brévignon, Christian Author Casagrande, Mirna M. Author Fåhraeus, Christer Author Grishin, Nick Author Kawahara, Akito Y. Author Mielke, Olaf H. H. Author Miller, Jacqueline Y. Author Nakamura, Ichiro Author Navas, Vanessa Author Patrusky, Brooke Author Pyrcz, Tomasz W. Author Richards, Lindsay Author Tan, Denise Author Tyler, Stephanie Author Viloria, Angel Author Warren, Andrew D. Author Xiao, Lei Author Freitas, André V. L. Author Willmott, Keith R. text Zoological Research 2023 2023-02-15 2023 1 73 journal article 58039 10.1111/syen.12590 bfb878f3-8a74-46d3-a104-36485c32aaba 7909395

TABLE 1 Comparison of characters for distinguishing Deltaya gen.n., Modica gen.n. and related genera.

	Modica gen.n.	Emeryus	Paryphthimoides	Colombeia	Scriptor	Deltaya gen.n.	Malaveria
Eyes: hair-like setae	Present	Absent	Present	Present	Present	Present	Present
DHW pale pupil dots in ocellus in cell Cu2-Cu1	Visible	Not visible	Variably visible	Variably visible	Not visible	Not visible	Not visible
VFW: prominent dark brown band (umbra) underlying postdiscal ocelli	Yes	No	Yes, somewhat (except almost absent in P. poltys, P. vestigiata)	Yes	Yes	Yes	No or weak umbra
VHW: dark marginal line in tornus	Thin, not broadening	Thin, not broadening	Thin, not broadening (except P. sheba, P. pseudoconfusa)	Marginal line slightly broader throughout wing	Broadening	Broadening	Thin, not broadening
VHW: postdiscal ocelli in cells Cu1- M3 and M3-M2	Double pupils (silver dots or elongate dashes) distinctly ringed with yellow	Where present, double pupils (silver dots) distinctly ringed with yellow	Variable across species, either double pupils (silver dots) distinctly ringed with yellow, or single pupil (an elongate silver smudge) indistinctly ringed with orange	Where present, double pupils (silver dots) distinctly ringed with yellow	Single pupil (an elongate silver smudge) indistinctly ringed with orange	Double pupils (silver dots or elongate dashes) distinctly ringed with yellow	Double pupils (silver dots or elongate dashes) distinctly ringed with yellow
Male genitalia aedeagus: cornuti	Variably present	Present	Present	Absent	Absent	Present (except in D. andrei and D. probata)	Absent
Female genitalia: lamella antevaginalis	No sclerotized lamella antevaginalis	Wrinkled, sclerotized lamella antevaginalis	Sclerotized lamella antevaginalis in some species	Sclerotized lamella antevaginalis	No sclerotized lamella antevaginalis	Sclerotized lamella antevaginalis (except in D. andrei and D. probata)	Sclerotized ‘spike’-like lamella antevaginalis