Hill forest dweller: a new cryptic species of Rhinolophus in the ‘ pusillus group’ (Chiroptera: Rhinolophidae) from Thailand and Lao PDR Author Soisook, Pipat Author Karapan, Sunate Author Srikrachang, Mattana Author Dejtaradol, Ariya Author Nualcharoen, Kwan Author Bumrungsri, Sara Author Lin, Sai Sein Author Aung, Moe Moe Author Bates, Paul J. J. Author Harutyunyan, Margarita Author Buś, Magdalena M. Author Bogdanowicz, Iesław text Acta Chiropterologica 2016 2016-12-31 18 1 117 139 journal article 22934 10.3161/15081109ACC2016.18.1.005 c723b052-3bef-46d2-903b-048ce2cebb1f 1508-1109 3756730

TABLE 1. Body mass (MASS; in grams), frequency of maximum energy (fmaxe; in kHz) and external measurements (in mm) of R. monticolus sp. nov., R. shortridgei, R. lepidus, R. refulgens and R. pusillus. Mean ± SD, min-max values are given. A sample size that differs from the total number of specimens is given in brackets. Definitions of measurement are included in the Materials and Methods section

n (sex)	MASS	fmaxe	FA	E	NL	HB	T	TIB	HF	5MET	4MET	3MET	3D1PH	3D2PH
R. monticolus sp. nov. (Thailand)
Holotype ♂	8.0	83.6	44.1	17.9	8.6	48.3	22.8	18.4	8.6	34.8	34.9	33.9	14.1	19.9
4 ♂♂	7.5 ± 0.8	85.9 ± 0.8	41.9 ± 0.48	16.8 ± 1.7	8.5, 8.6 [2]	46.5 ± 2.6	23.0 ± 3.0	17.7 ± 1.0	7.8 ± 0.4	33.2 ± 1.2	33.2 ± 1.3	32.4 ± 1.7	12.7 ± 0.4	18.0 ± 0.8
	6.9–8.6	85.2–87.0	41.4–42.3	15.7–19.4	–	42.7–48.5	19.7–25.6	16.7–18.6	7.4–8.3	32.0–34.7	32.0–35.1	31.2–34.9	12.2–13.1	16.9–18.7
2 ♀♀	7.2 [1]	91.6, 93.0	41.2, 42.2	15.5, 15.8	7.1 [1]	42.3, 45.2	22.1, 24.8	16.9, 17.5	7.4, 8.0	31.9, 33.0	32.3, 34.1	31.3, 32.8	12.2, 12.9	16.3, 18.0
R. shortridgei (Type locality, Bagan, Myanmar)
Holotype ♂	–	–	41.0	–	–	–	17.3	18.2	8.1	–	–	–	–	–
7 ♂♂	7.4 [1]	94.5 [1]	40.7 ± 1.4	15.5 [1]	7.9 [1]	45.4 [1]	20.7 ± 2.7	17.1 ± 1.04	8.3 ± 1.0	33.8 [1]	34.3 [1]	33.9 [1]	12.6 [1]	17.2 [1]
	–	–	38.7–41.9	–	–	–	16.1–24.2 [6]	15.8–18.4 [6]	6.0–9.7 [6]	–	–	–	–	–
2 ♀♀	7.0 [1]	98.8 [1]	40.4, 41.7	15.1 [1]	6.9 [1]	47.7 [1]	20.2, 23.3	17.7, 18.0	8.7, 9.1	33.8 [1]	34.1 [1]	33.0 [1]	12.6 [1]	17.0 [1]
R. shortridgei (elsewhere in Myanmar)
9 ♂♂	7.1 ± 1.0	96.8 ± 1.0	40.3 ± 1.4	16.0 ± 1.7	6.7, 6.9 [2]	43.3 ± 2.2	19.8 ± 2.5	17.4 ± 1.2	8.4 ± 0.5	32.3 ± 1.5	32.6 ± 1.2	31.2 ± 1.1	11.9 ± 0.5	17.5 ± 1.6
	6.0–8.8 [8]	95.9–97.9 [3]	38.2–42.1	14.0–18.7	–	40.6–47.8	16.2–25.0	15.6–19.0	7.9–9.4	29.8–34.0	29.9–34.0	29.1–32.3	11.1–12.7	15.3–19.9
2 ♀♀	5.5–6.2	100.7 [1]	39.4, 40.2	16.7, 17.9	6.9 [1]	43.8, 46.4	21.1, 29.4	16.0, 17.6	6.9, 8.3	31.9, 32.8	32.2, 32.2	30.4, 30.9	11.4, 12.1	16.8, 18.0
R. lepidus (India)
5 ♂♂	4.0, 5.0 [2]	104.0, 105.0 [2]	40.5 ± 1.6	12.6 ± 0.7	–	40.6 ± 2.0	17.7 ± 3.4	16.3 ± 0.3	7.0 ± 0.7	31.5 ± 0.6	31.3 ± 1.2	31.3 ± 0.9	11.4 ± 0.4	17.4 ± 0.6
	–	–	38.7–42.2 [4]	11.8–13.5 [4]	–	37.7–43.0	13.3–21.0	16.1–16.6 [4]	6.2–7.6	31.1–32.2 [3]	30.4–32.6 [3]	30.4–32.2 [3]	11.0–11.6 [3]	16.8–18.0 [3]
7 ♀♀	5.00	–	41.1 ± 1.0	14.5 ± 1.6	–	41.4 ± 7.6	19.1 ± 3.8	16.1 ± 0.7	7.8 ± 0.9	31.6 ± 0.9	31.7 ± 1.1	31.3 ± 1.7	11.1 ± 0.5	18.1 ± 0.6
	–	–	40.0–42.9 [6]	13.1–16.8 [6]	–	33.5–52.0 [5]	14.0–23.0 [5]	15.1–17.2 [6]	7.2–9.7	30.2–32.2 [5]	30.2–33.1 [5]	29.6–33.7 [5]	10.2–11.5 [5]	17.1–18.5 [5]
R. lepidus (Myanmar, N. Thailand and Vietnam)
4 ♂♂	–	–	38.9 ± 0.9	15.5 ± 1.6	–	41.1 ±1.4	18.2 ± 2.2	16.0 ± 0.4	6.6 ± 0.6	30.1 ± 0.5	30.2 ± 0.8	28.9 ± 1.2	10.8 ± 0.6	16.4 ± 1.3
	–	–	37.7–39.9	13.3–17.3	–	39.7–43.0	15.0–19.8	15.8–16.5	5.7–7.1	29.7–30.8	29.1–31.2	27.8–30.2	10.0–11.2	15.1–17.7
2 ♀♀	6.0 [1]	103.0 [1]	39.7, 40.2	15.3, 16.6	–	38.2, 41.9	19.8, 20.6	16.1, 17.1	6.9, 7.5	29.8, 31.0	30.1, 31.5	28.7, 30.0	10.8, 12.4	16.8, 17.1
R. refulgens (S. Thailand, peninsular Malaysia)
12 ♂♂	5.6 ± 1.0	102.2 ± 2.1	39.4 ± 1.5	15.0 ± 1.5	–	45.7 ± 3.3	19.4 ± 2.0	16.4 ± 0.9	7.2 ± 0.4	29.3 ± 1.4	29.8 ± 1.3	28.7 ± 1.2	10.8 ± 0.6	15.7 ± 1.2
	4.0–7.2 [11]	98.6–105.2	36.6–42.6	12.3–17.8	–	39.0–51.1	16.2–22.4	14.6–18.0	6.3–7.9	26.0–31.7	26.9–32.2	25.7–30.7	9.5–11.4	13.1–17.6
10 ♀♀	4.8 ± 0.5	102.0 ± 1.3	38.8 ± 1.0	15.1 ± 1.1	–	41.3 ± 3.8	19.2 ± 2.2	15.7 ± 0.5	6.7 ± 0.7	28.9 ± 0.6	29.7 ± 0.6	28.6 ± 0.5	10.6 ± 0.5	15.0 ± 1.0
	4.0–5.5, [9]	100.7–104.1	37.2–40.4	13.6–16.8	–	37.2–49.0	14.6–21.6	14.9–16.5	5.3–7.6	28.2–30.2	28.6–30.3	27.9–29.2	10.0–11.3	13.9–16.6
R. pusillus (Thailand)
25 ♂♂	4.7 ± 0.8	111.1 ± 2.0	37.1 ± 1.0	14.4 ± 1.3	–	38.4 ± 4.6	17.5 ± 2.2	15.7 ± 0.7	6.3 ± 0.6	27.8 ± 0.8	28.2 ± 0.7	27.1 ± 0.8	10.3 ± 0.4	14.1 ± 0.7
	3.0–7.0 [22]	108.4–115.2 [22]	34.9–38.9	11.4–17.3	–	29.8–45.5	13.4–21.3	14.7–17.1	4.0–7.1	26.1–29.5	26.7 ± 29.5	25.6–28.6	9.3–11.2	12.6–15.3
18 ♀♀	4.6 ± 0.5	110.4 ± 1.8	37.4 ± 1.1	13.9 ± 1.1	–	37.5 ± 3.3	17.7 ± 1.7	15.6 ± 0.8	6.5 ± 0.5	28.1 ± 0.9	28.4 ± 1.0	27.4 ± 0.8	10.4 ± 0.4	14.4 ± 0.8
	4.0–5.5 [13]	107.3–113.1	35.8–39.4	10.8–16.0	–	32.3–42.6	13.7–21.1	14.5–17.4	5.6–7.3	25.9–29.6	26.5–30.3	25.9–29.0	9.8–11.1	13.0–16.2