Hill forest dweller: a new cryptic species of Rhinolophus in the ‘ pusillus group’ (Chiroptera: Rhinolophidae) from Thailand and Lao PDR Author Soisook, Pipat Author Karapan, Sunate Author Srikrachang, Mattana Author Dejtaradol, Ariya Author Nualcharoen, Kwan Author Bumrungsri, Sara Author Lin, Sai Sein Author Aung, Moe Moe Author Bates, Paul J. J. Author Harutyunyan, Margarita Author Buś, Magdalena M. Author Bogdanowicz, Iesław text Acta Chiropterologica 2016 2016-12-31 18 1 117 139 journal article 22934 10.3161/15081109ACC2016.18.1.005 c723b052-3bef-46d2-903b-048ce2cebb1f 1508-1109 3756730

TABLE 3. Matrix summarising discriminating features of Rhinolophus in the ‘ pusillus group’ currently recognised from mainland Southeast Asia. Characters that are most distinct from R. monticolus sp. nov. are highlighted in grey

Species	fmaxe (kHz)	Forearm length (FA; mm)	Skull length (SL; mm)	Sella and lancet shape	Posterior median chambers of rostrum	Supraorbital region	Upper canine (C 1)	Second lower premolar (P 3)	Third lower premolar (P 4)
R. monticolus	85.2–93.0	41.2–44.1	17.44–19.71	Sella broad, parallel sided or slightly convex in the middle; squared-off tip. Lancet tall; slightly concave sides; bluntly pointed tip	Inflated; well defined	Narrow and shallow; supraorbital ridges poorly developed	Twice P 4 in height equal or only slightly exceeding P 4 in crown area	Rounded and very small; in the main axis or slightly extruded from the toothrow	Rounded; equal to P 2 in crown area
R. shortridgei	94.5–100.7	38.2–41.0	17.17–18.60	Sella broad, parallel sided; rounded tip. Lancet tall; slightly concave sides; bluntly pointed tip	Not inflated; poorly defined	Broad and deep; supraorbital ridges well developed	Twice P 4 in height; exceeding P 4 in crown area	Rounded and very small; half-way or extruded from the toothrow	Elongated; exceeds P 2 in crown area
R. lepidus	103.0–105.0	37.7–42.9	15.74–17.27	Sella narrow, parallel sided; rounded tip. Lancet tall; slightly concave sides; bluntly pointed tip	Slightly inflated; well defined	Narrow and deep; supraorbital ridges well developed	Twice P 4 in height; exceeding P 4 in crown area	Rounded and very small; slightly extruded from the toothrow	Rounded; exceeds P 2 in crown area
R. refulgens	98.6–105.2	36.6–42.6	15.75–17.10	Sella narrow, parallel sided or slightly convex; rounded or squared-off tip. Lancet tall; slightly concave sides; bluntly pointed tip	Slightly inflated; well defined	Broad and shallow; supraorbital ridges poorly developed	Slightly exceeds P 4 in height and crown area	Rounded and very small; usually, but not always extruded from the toothrow	Rounded; exceeds P 2 in crown area
R. pusillus	107.3–115.2	35.8–39.4	14.93–16.09	Sella narrow, parallel sided; squared-off tip. Lancet tall; slightly concave sides; bluntly pointed tip	Inflated; well defined	Narrow and shallow; supraorbital ridges well developed	Only slightly exceeds P 4 in height and crown area	Rounded and very small; usually extruded from the toothrow	Rounded; exceeds P 2 in crown area
R. acuminatus	85.4–94.3	45.7–50.0	19.02–22.21	Sella narrow, parallel sided; rounded tip. Lancet very short; narrow pointed tip	Not inflated; poorly defined	Broad and deep; supraorbital ridges well developed	Exceeds P 4 in height and crown area	Rounded and very small; usually extruded from the toothrow	Rectangular; exceeds P 2 in crown area