A new genus and tribal placement for Ceneus speculiferus Fairmaire, 1879, comments on Grundmannius Basilewsky, 1965, Mirachlaenius Facchini, 2011 and Stenoodes Basilewsky, 1953, and a key to the Oriental and Australasian genera of Chlaeniina Brullé, 1834 (Coleoptera, Carabidae) Author Guéorguiev, Borislav National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia (Bulgaria) gueorguiev@nmnhs.com Author Will, Kipling University of California Berkeley, Essig Museum of Entomology, 1101 VLSB # 4780, Berkeley, CA 94720 (United States) kipwill@berkeley.edu text Zoosystema 2025 2025-05-23 47 15 231 259 https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2025v47a15.pdf journal article 310029 10.5252/zoosystema2025v47a15 8d4b76e7-28ef-4620-bdcc-f494df80b527 1638-9387 15502564 11740D9A-34E8-48B0-A6C9-15AB438BA100
APPENDIX 1. — List of species and subspecies and number of specimens studied, with data for their provenance and repository.
Species and subspeciescountries specimens collections
Callistus lunatus (Fabricius, 1775)Bulgaria, Turkey (Koru Dağları)3NMNHS
Chlaenius (Achlaenius) variicornis A. Morawitz, 1863North Korea (Mount Myohyang)1coll. EK
Chlaenius (Amblygenius) anthraceus Alluaud, 1918Nigeria1NMNHS
Chlaenius (Amblygenius) episcopalis Dejean, 1831 (s.l.)Zimbabwe (Victoria Falls)1NMNHS
Chlaenius (Amblygenius) glabratus Dejean, 1831Sudan (South Darfur)3NMNHS
Chlaenius (Amblygenius) lioderus Andrewes, 1923India (Delhi)1NMNHS
Chlaenius (Amblygenius) liothorax Alluaud, 1934Nigeria1NMNHS
Chlaenius (Amblygenius) macropus Chaudoir, 1876Nepal (Lumbini)1coll. EK
Chlaenius (Amblygenius) omochlorus Andrewes, 1931Nepal (Trisuli Bazar)1NMNHS
Chlaenius (Amblygenius) pretiosus Chaudoir, 1856India (Delhi)4NMNHS
Chlaenius (Amblygenius) rufomarginatus Dejean, 1831Zimbabwe (Victoria Falls)1NMNHS
Chlaenius (Amblygenius) trachys Andrewes, 1923Sri Lanka (Dambulla)1NMNHS
Chlaenius (Amblygenius) sp.Nepal (Lumbini), Sri Lanka (Dambyla)2NMNHS
Chlaenius (Callistoides) amabilis Chaudoir, 1876Nepal (Lumbini)3NMNHS
Chlaenius (Callistoides) malachinus (Motschulsky, 1865)India (Delhi)1NMNHS
Chlaenius (Callistoides) tripustulatus (Dejean, 1831)Nigeria3NMNHS
Chlaenius (Callistoides) sp.Sri Lanka (Dambyla)1NMNHS
Chlaenius (Capsochlaenius) luisae Gestro, 1895Somalia2coll. EK
Chlaenius (Chlaeniellus) atratulus Mandl, 1983 (s.l.)India (Ladakh)1NMNHS
Chlaenius (Chlaeniellus) extensus Mannerheim, 1825Turkmenistan2NMNHS
Chlaenius (Chlaeniellus) flavipes Ménétriés, 1832Albania, Bulgaria, Turkey8NMNHS
Chlaenius (Chlaeniellus) impressicollis Chaudoir, 1876India (Elephanta Isl.)1NMNHS
Chlaenius (Chlaeniellus) koenigi Semenov, 1888Turkmenistan1coll. EK
Chlaenius (Chlaeniellus) nigricornis (Fabricius, 1787)Bulgaria3NMNHS
Chlaenius (Chlaeniellus) semiviridis Andrewes, 1920Indonesia (W Timor)4NMNHS
Chlaenius (Chlaeniellus) stenoristus Chaudoir, 1876India (Delhi)1NMNHS
Chlaenius (Chlaeniellus) terminatus Dejean, 1826Bulgaria1NMNHS
Chlaenius (Chlaeniellus) tristis tristis (Schaller, 1783)Bulgaria2NMNHS
Chlaenius (Chlaeniellus) vestitus (Paykull, 1790)Albania, Bulgaria, Greece, Turkey, Armenia53NMNHS
Chlaenius (Chlaeniellus) sp.Thailand1NMNHS
Chlaenius (Chlaeniodromus) tetraphacus Alluaud, 1918Ivory Coast2coll. EK
Chlaenius (Chlaeniostenus) angustatus Dejean, 1831Nigeria1NMNHS
Chlaenius (Chlaeniostenus) circumdatus Brullé, 1835 (s.l.)Nepal (Lumbini), Sri Lanka (Polonnaruwa), Indonesia (Mentawan Isl.)5NMNHS
Chlaenius (Chlaeniostenus) nitidicollis Dejean, 1826India (Delhi)2NMNHS
Chlaenius (Chlaeniostenus) sulcipennis sulcatulus Boheman, 1848Mozambique (Cabo Delgado)1NMNHS
Chlaenius (Chlaenites) spoliatus spoliatus (P. Rossi, 1792)Bulgaria5NMNHS
Chlaenius (Chlaenius) dusaultii (L. Dufour, 1821) (s.l.)Sudan (South Darfur & West Darfur), Nigeria8NMNHS
Chlaenius (Chlaenius) festivus festivus (Panzer, 1796)Albania, North Macedonia, Bulgaria, Greece, Armenia, Iraq18NMNHS
Chlaenius (Chlaenius) festivus velutinus (Duftschmid, 1812)Spain3NMNHS
Chlaenius (Chlaenius) meticulosus Laferté-Sénectère, 1851Senegal, Nigeria2coll. EK
Chlaenius (Chlaenius) pseudocruciatus Kirschenhofer, 2009Sudan (South Darfur)3NMNHS, coll. EK
Chlaenius (Dinodes) cruralis Fischer von Waldheim, 1829North Macedonia1NMNHS
Chlaenius (Dinodes) decipiens (L. Dufour, 1820)North Macedonia, Bulgaria, Greece17NMNHS
Chlaenius (Epomis) dejeanii (Dejean, 1831)Turkey (Adana)1NMNHS
Chlaenius (Epomis) duvaucelii (Dejean, 1831)India (Delhi)1coll. EK
Chlaenius (Epomis) nigricans Wiedemann, 1821India (Dehradun, Delhi), Nepal (Lumbini), Sri Lanka (Polonnaruwa), Indonesia (Lombok)8NMNHS, coll. EK
Chlaenius (Haplochlaenius) baehri (Kirschenhofer, 1998)Indonesia (Sulawesi)1SMNS
Chlaenius (Haplochlaenius) costiger costiger Chaudoir, 1856North Korea (Kumgangsan)1coll. EK
Chlaenius (Iberodinodes) dives Dejean, 1826 (s.l.)Spain1NMNHS
Chlaenius (Lissauchenius) borgouensis Kirschenhofer, 2007Nigeria2NMNHS
Chlaenius (Lissauchenius) darfurensis Kirschenhofer, 2007Sudan (West Darfur)1coll. EK
Chlaenius (Lissauchenius) rufifemoratus lynx Chaudoir, 1856North Korea (Suyangsan Falls)1coll. EK
Chlaenius (Nectochlaenius) canariensis seminitidus Chaudoir, 1856Israel1coll. EK
Chlaenius (Nectochlaenius) persicus L. Redtenbacher, 1850Iran (Shiraz)1coll. EK
Chlaenius (Ocybatus) dyrollei (Laferté-Sénectère, 1851) (s.l.)Nigeria2NMNHS
Chlaenius (Ocybatus) discicollis (Laferté-Sénectère, 1851)Nigeria2NMNHS
Chlaenius (Pachydinodes) bipustulatus Boheman, 1848Nigeria1NMNHS
Chlaenius (Pachydinodes) pictus Chaudoir, 1856India (Delhi)1NMNHS
Chlaenius (Pachydinodes) posticus (Fabricius, 1798)Sri Lanka (Polonnaruwa)4NMNHS
Chlaenius (Pachydinodes) virgulifer Chaudoir, 1876India (Delhi)1coll. EK
Chlaenius (Pachydinodes) sp.Sudan (South Darfur)1NMNHS
Chlaenius (Pleroticus) buquetii hacquardi Chaudoir, 1883Zaire (Kisangani)1NMNHS
Chlaenius (Prochlaeniellus) senegalensis Dejean, 1831Sudan (South Darfur), Nigeria2NMNHS
Chlaenius (Pseudochlaeniellus) chitwanensis Kirschenhofer, 2005 Nepal (Lumbini)1NMNHS
Chlaenius (Rhopalopalous) janthinus L.Redtenbacher, 1844India (Jammu & Kashmir)5NHMUK
Chlaenius (Rhysotrachelus) nepalensis Hope, 1831Nepal (Betrawati)1NMNHS
Chlaenius (Sphodromimus) holzschuhi Casale, 1984Nepal1SMNS
Chlaenius (Stenochlaenius) coeruleus (Steven, 1809)Turkey (Artvin)4NMNHS
Species and subspeciescountriesspecimens collections
Chlaenius (Tomochilus) alternans (Imhoff, 1843) (s.l.)Nigeria2NMNHS
Chlaenius (Tomochilus) carbonatus Chaudoir, 1876Senegal, Burkina Faso, Nigeria, Ethiopia5NMNHS, coll. EK
Chlaenius (Trichochlaenius) aeneocephalus aeneocephalus Dejean, 1826Bulgaria10NMNHS
Chlaenius (unplaced) rufithorax Wiedemann, 1821India2NHMUK
Grundmannius dispar (Péringuey, 1896)Mozambique, South Africa4CAS, NMNHS
Harpaglossus laevigatus (Dejean, 1828)“W. Africa”, Senegal3NHMUK, SMNS
Harpaglossus opacus (Chaudoir, 1857)Eritrea, Saudi Arabia, India7IZAS, NHMUK
Hololeius ceylanicus (Nietner, 1856)Sri Lanka, Myanmar, Indonesia (Bali), Australia19NMNHS, NHMUK, SMNS
Holosoma hedini (Andrewes, 1935)China (Gansu)2NMNHS
Holosoma weigoldi (Heller, 1923)China (Sichuan)3NMNHS
Lapitachlaenius speculiferus (Fairmaire, 1879) n. comb.Fiji17ANIC, BPBM, NZAC, SAMA
Lapita chlaenius specularis (Emden, 1937) n. comb.Vanuatu3NHMUK
Mirachlaenius barbarae Facchini, 2011India (Nilgiri Hills)1SMNS
Parachlaenius (Parachlaenius) eminus Kolbe, 1894Uganda, Tanzania (Bukoba)3NHMUK, SMNS
Parachlaenius (Parachlaenius) violaceus Péringuey, 1899“E. Africa”, Zimbabwe5NHMUK, SMNS
Procletodema parallelum Péringuey, 1899Zimbabwe3NHMUK, SMNS
Procletus minor Basilewsky, 1968Nigeria (Samaru)1NHMUK
Procletus singularis Péringuey, 1896Zimbabwe3NHMUK, SMNS
Rhopalomelus angusticollis Boheman, 1848South Africa6NHMUK, SMNS
Stenoodes jeanneli Basilewsky, 1953Madagascar1MNHN
Stuhlmannium mirabile Kolbe, 1894“E. Africa”1NHMUK