A new limestone-associated species of wolf snake in the Lycodon fasciatus complex (Squamata: Serpentes: Colubridae) from central Vietnam Author Vogel, Gernot 0000-0002-4542-518X Society for South East Asian Herpetology, Im Sand- 3, D- 69115 Heidelberg, Germany ogel@t-online.de Author Bragin, Andrey M. 0000-0002-3621-9763 Joint Vietnam-Russia Tropical Science and Technology Research Centre, Nghia Do, Cau Giay, Hanoi, Vietnam bragin98@yandex.ru Author Poyarkov, Nikolay A. 0000-0002-3621-9763 Joint Vietnam-Russia Tropical Science and Technology Research Centre, Nghia Do, Cau Giay, Hanoi, Vietnam & Department of Vertebrate Zoology, Lomonosov Moscow State University, Leninskiye Gory, GSP- 1, Moscow 119991, Russia bragin98@yandex.ru Author Nguyen, Tan Van 0000-0001-5413-968X Institute for Research and Training in Medicine, Biology and Pharmacy, Duy Tan University, Da Nang, 550000, Vietnam tan.sifasv@gmail.com text Zootaxa 2025 2025-08-12 5679 1 1 44 https://doi.org/10.11646/zootaxa.5679.1.1 journal article 312182 10.11646/zootaxa.5679.1.1 688f360d-42eb-4a61-9421-df275e9723a6 1175-5326 16983543 B743F819-2EFE-48EC-A561-02ABC6D97947

TABLE 3. Comparison of morphological characters of Lycodon calcarophilus sp. nov. with those of the Lycodon fasciatus group. Characters highlighted in bold are considered diagnostic (with respect to Lycodon calcarophilus sp. nov.). Numeric values provided for SVL refer to the maximum SVL attained by each species.

Species	Max SVL (mm)	TaL/TL	VEN	SC	BB	TB	DSR keeled?	CP	LOR-E	Dorsal ground color	Color of light crossbands	Ventral pattern	Distributions	Sources
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calcarophilus sp. nov.	505	0.19–0.22	214–219	83–88	19–25	10–15	Keeled	Entire	Yes	Black	White anteriorly, slight brownish-grey posteriorly	Banded	Vietnam	This study
butleri	648	0.19–0.24	216–228	81–97	28–37	14–25	Keeled	Entire	Yes	Grey or black	White or grey	Banded & spotted	Thailand, Peninsular Malaysia	Grismer et al. (2014); Nguyen et al. (2024a)
davidi	308	0.21	224	99	84	38	Keeled	Entire	No	Dark brown	Brownish-grey	Banded & spotted	Laos	Vogel et al. (2012)
cavernicolus	406	0.20–0.21	232–245	92–113	36–45	29–41	Keeled	Entire	Yes	Light brown	White or light brown	Plain	Thailand; Peninsular Malaysia	Grismer et al. (2014); Vogel et al. (2024)
fasciatus	706	0.19–0.22	197–213	74–90	28–43	10–20	Keeled	Entire	Yes	Black or dark brown	White or light brown anteriorly, variable posteriorly, usually light brown or reddish brown	Banded & spotted	China, Myanmar, India, Bhutan, Thailand, Laos, Vietnam	Nguyen et al. (2024a); Vogel et al. (2024)
gammiei	1110	0.19–0.25	205–228	101–112	42–58	16–23	Keeled	Entire	No	Black	Yellow or olive yellow	Banded	India, Bhutan, China	Mistry et al. (2007); Chettri & Bhupathy (2009); Shu et al. (2024)
gongshan	798	0.21–0.27	203–216	79–96	26–46	9–16	Keeled	Entire	Yes	Black	White anteriorly, variable posteriorly, usually orange-brown or reddish-brown	Variable (banded or banded & spotted)	China, Myanmar	Vogel & Luo (2011); Huang et al. (2021); Nguyen et al. (2024a)
latifasciatus	680	0.21–0.23	206–216	90–96	29–36	13–21	Keeled	Entire	Yes (No)	Dark brown or black	Orange or light brown	Banded & spotted	China, Myanmar	Nguyen et al. (2025)
liuchengchaoi	676	0.16–0.22	200–228	68–81	18–54	7–16	Keeled	Divided or Entire	Yes	Black	Cream anteriorly, yellow, orange or light brown posteriorly	Banded	China	Zhang et al. (2011); Nguyen et al. (2024a)
multifasciatus	570	0. 23–0.26	227–237	106–117	51–72	25–35	Keeled	Entire	No	Dark brown	Light brown	Plain	Japan	Maki (1931); Vogel et al. (2009); Nguyen et al. (2024a)
multizonatus	505	0. 16–0.18	191–202	56–75	55–62	11–19	Keeled	Divided	Yes	Black	Orange or reddishbrown	Variable (plain or banded)	China	Lei et al. (2014); Nguyen et al. (2024a)
obvelatus	447	0.19	199	76	31	13	Smooth	Entire	Yes	Black	Light orange or salmon	Banded & spotted	China	Wang et al. (2021); Nguyen et al. (2024a)
pictus	543	0.18–0.23	209–218	82–91	25–32	8–16	Smooth	Entire	Yes or No	Black or dark brown	White or cream anteriorly, light brown posteriorly	Banded	Vietnam, China	Janssen et al. (2019, 2020); Xu et al. (2023); Nguyen et al. (2024a)
ruhstrati abditus	762	0.19–0.25	210–230	90–115	17–47	9–23	Keeled	Entire	No	Dark brown	White anteriorly, white or grey-brown posteriorly	Plain & spotted	China, Vietnam, Laos	This study
ruhstrati ruhstrati	810	0.21–0.25	211–228	99–114	33–46	14–25	Keeled	Entire	No	Black or dark brown	Grey-brown	Plain & spotted	Taiwan Island	This study
serratus	480	0. 24	198	84	66	26	Smooth	Divided	Yes	Dark brown	Olive brown (tawny olive)	Banded & spotted	China	Wang et al. (2021)
sidiki	650	0.19–0.24	195–212	77–91	25–33	10–16	Keeled	Entire	Yes	Black	Light grey	Banded	Indonesia (Sumatra)	Wostl et al. (2017); Nguyen et al. (2024a)
synaptor	395	0.19	201–203	68–69	30–31	9	Keeled	Entire	No	Black or dark brown	White	Banded	China	Vogel & David (2010)
yunnanensis	550	0.18–0.21	186–202	67–80	20–36	7–12	Keeled	Entire	Yes	Black or dark brown	White	Banded & spotted	China, Vietnam	Vogel et al. (2024)