High-level taxonomic splitting in allopatric taxa causes confusion downstream: a revision of the nudibranch family Сoryphellidae Author Ekimova, Irina Author Valdés, Ángel Author Malaquias, Manuel António E Author Rauch, Cessa Author Chichvarkhin, Anton Author Mikhlina, Anna Author Antokhina, Tatiana Author Chichvarkhina, Olga Author Schepetov, Dimitry text Zoological Journal of the Linnean Society 2022 2022-09-01 196 1 215 249 https://academic.oup.com/zoolinnean/article/196/1/215/6549318 journal article 133193 10.1093/zoolinnean/zlab109 85c62267-04fb-4e36-8c00-1ef4bb11c343 0024-4082 7031383

Table 4. Variability of morphological characters in the target species

Species	Coryphella verrucosa	Coryphella	Borealea nobilis	Borealea	Microchlamylla	Microchlamylla
	(M. Sars, 1829)	pseudoverrucosa	(Verrill, 1880)	sanamyanae	gracilis (Alder &	amabilis (Hirano &
		Martynov et al.,		(Korshunova et al.,	Hancock, 1844)	Kuzirian, 1991)
		2015		2017)
Body	Narrow	Narrow	Wide	Wide	Narrow	Narrow
Notal ridge	Completely reduced	Completely reduced	Reduced, continuous	Reduced,	Reduced, discon-	Reduced,
				continuous	tinuous	discontinuous
Cerata	Several groups;	Several groups; long	Continuous rows	Continuous rows	Several groups;	Several groups; long
	‘verrucosa’ morph	cerata			long cerata	cerata
with short cerata and
‘rufibranchialis’ morph
with long cerata
Rhinophores	Smooth with small tuber-	Smooth sometimes	Wrinkled	Smooth, rarely	Smooth	Smooth
	cles	with small tuber-		slightly wrinkled
		cles
Coloration	Variable: cerata from	Cerata from in-	Cerata from beige	Cerata from	Cerata from orange	Cerata intensive red
	pale beige to intensive	tensive red to	to pink, reddish-	reddish-orange	to intensive red	or reddish-orange;
	red, wine or brown,	reddish-brown,	orange and bright	to bright red;	or reddish-brown;	rhinophores white
	sometimes with nu-	sometimes with	orange; rhinophores	rhinophores	rhinophores
	merous white dots;	numerous white	orange to light pink	white with white	white
	rhinophores from white	dots; rhinophores	with white pigment	opaque pigment
	to pale orange or pink	white	on top	covered 2/3 of
				their length
Jaws masticatory	From 1 to 7 rows of	From 1 to 7 rows of	From 9 to 12 rows of	From 3 to 6 rows	From 4 to 6 rows	From 4 to 6 rows of
process	denticles	denticles	strong conical dent-	of strong conical	of small conical	small conical dent-
			icles	denticles	denticles	icles
Rachidian teeth	Triangle plate with non-	Triangle plate with	Triangle plate of	Triangle plate of	Triangle plate with	Triangle plate with
	compressed conical	non-compressed	different propor-	different pro-	non-compressed	non-compressed
	cusp of different pro-	conical cusp of dif-	tions with non-	portions with	elongated con-	elongated conical
	portions and 5–9 dent-	ferent proportions	compressed or	non-compressed	ical cusp and 6–9	cusp and 7–9 dent-
	icles on each side	and 5–8 denticles	slightly compressed	or slightly com-	denticles on each	icles on each side
		on each side	conical cusp and	pressed conical	side forming deep	forming deep fur-
			7–9 denticles on	cusp and 6–8	furrows	rows
			each side	denticles on each
				side
Lateral teeth	Triangle, smooth or with	Triangle, bearing	Triangle, bearing	Triangle, bearing	Triangle, bearing	Triangle, bearing 5–7
	7–12 small denticles,	8–13 small dent-	10–15 small dent-	8–11 small dent-	7–9 small dent-	small denticles,
	without attenuated	icles, without at-	icles, with at-	icles, with at-	icles, with at-	with attenuated
	basally process	tenuated basally	tenuated basally	tenuated basally	tenuated basally	basally process
		process	process	process	process