New records on the rich loriciferan fauna of Trezen ar Skoden (Roscoff, France): Description of two new species of Nanaloricus and the new genus Scutiloricus Author Neves, Ricardo Cardoso Author Kristensen, Reinhardt Møbjerg Natural History Museum of Denmark, University of Copenhagen, Copenhagen Ø, Denmark Author Møbjerg, Nadja Department of Biology, August Krogh Building, University of Copenhagen, Copenhagen Ø, Denmark, text PLoS ONE 2021 e 0250403 2021-05-05 16 5 1 42 journal article 301517 10.1371/journal.pone.0250403 b83f9ed2-1a72-4f1e-b9b0-10d6940f37fa 1932-6203 PMC8099068 33951070 13151726

Table 1. Comparison of adult characters of the species assigned to the genus Nanaloricus (family Nanaloricidae). Nanaloricus khaitatus is not included in this table because its original description is probably based on observations of at least three different species.

		N. mysticus [8, 15]	N. gwenae [30]	N. valdemari nov. sp. (this study)	N. mathildeae nov. sp. (this study)
https://doi.org/10.1371/journal.pone.0250403.t001
Mouth	Length	short	long	long	long
cone:
	Honeycomb	absent	absent	present	absent
sculpture
	Oral ridges	8, of equal length	8, four primary oral	8, four primary oral ridges one-	8, four primary oral ridges one-
			ridges twice as long as the	fourth as long as four secondary	fourth as long as four secondary
			four secondary
Introvert:	Female	slender, with numerous papillae	slender, two-segmented,	slender, club-shaped, four-	slender, club-shaped, four-
	clavoscalids	at the base and a filiform tip	with several small, spike-	segmented, with many small	segmented, with several long
			like papillae at the base	papillae and a spinose tip	papillae and a spinose tip
			and a hook-shaped tip
	Male	multiform: midventral pair	?	multiform: midventral pair four-	multiform: midventral pair four-
	clavoscalids	slender; six clavoscalids ramified	(males were never found)	segmented, slender, with spinose	segmented, slender, with spinose tip;
		into three branches; the tertiary		tip; six clavoscalids ramified into	six clavoscalids ramified into three
		branches broad and club-shaped		three branches; the primary	broad, flat branches; the primary
		(except in the laterodorsal pair)		branch three-segmented, with	branch three-segmented, with
				spinose tip	spinose tip
	8 th row	unsegmented, whip-like	two-segmented, distal	unsegmented, whip-like	unsegmented, whip-like
	spinoscalids		segment with serrated
			margins
Neck:	Double and	all with blunt tips, uniform; in	all with blunt tips, but	all with blunt tips, but upper row	upper row appendages of the double
	single	males, the upper row	upper row appendages of	appendages of the double	trichoscalids have a pointy tip and
	trichoscalids	appendages of the lateroventral	the double trichoscalids	trichoscalids are longer	are longer than those in the lower
		pair is modified into claspers	are slightly thinner than	(ventrolateral pairs) or slightly	row, which have a blunt tip; single
		(claw-shaped organs)	lower row	longer (lateral pairs) than lower	trichoscalids are broader than the
				row	double trichocalids, and possess a
					unique blunt tip with several (ca.
					14?) finger-like processes
	Sensory organ	not found	not found (though a	present, located at the most	present, located anteroproximally to
	of the double		short, spine-like structure	anterior margin of the upper	the ventral and ventrolateral double
	trichoscalids		is present in the drawings	appendage basal plate of the	trichoscalids, and protrudes from its
			of this species	ventral and ventrolateral double	own plate with a double-square
			description)	trichoscalids	shape
Lorica:	Nr. of plates	6	6	6	6
	Anterior spikes	15 (midventral spike short)	15 (midventral spike	15 (midventral spike very short)	15 (midventral spike very short)
			short)
	Honeycomb	present	present	present	present
sculpture
	Number of N-	4 large, arranged rectangularly	4 large, arranged	4 large, arranged rectangularly on	4 large, arranged rectangularly on
	flosculi	on each dorsolateral plate; 1	rectangularly on each	each dorsolateral plate; 1 large,	each dorsolateral plate; 1 large,
		large, located posteriorly on the	dorsolateral plate; 1 large,	located posteriorly on the dorsal	located posteriorly on the dorsal
		dorsal plate	located posteriorly on the	plate	plate
			dorsal plate