| Colour | brownish-black | reddish brown to black | uniform orange-brown |
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| Cephalic/pronotal punctuation | weak, irregular | weak, irregular | dense and strong |
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| Clypeo-frontal tubercles | small, closely spaced | small, closely spaced | well developed, subconical, directed forward |
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| Mentum | transverse, the disc evenly convex (Fig. 15) | subtrapezoidal with basal fovea (Fig. 16) | distinctly transverse, subplanar (Fig. 17) |
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| Eyes | medium-sized (Fig. 11) | large, prominent (Fig. 12) | reduced (Fig. 13) |
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| Mandibles (♂) | subequal | subequal | asymmetrical, left mandible much more developed, directed upwards |
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| Elytra length | about 1.75 × pronotum length | nearly twice pronotum length | about 1.65 × pronotum length |
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| Striae | superficial | superficial | deeply impressed |
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| Aedeagus (parameres, frontal view) | outer margins of parameres evenly convex (Figs. 18, 19) | outer margins straight; parameres obliquely converging from base to apex (Figs. 21, 22) | parameres relatively short; outer margins parallel (Figs. 24, 25) |
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| Endophallites | Fig. 20 | Fig. 23 | Fig. 26 ( *) |
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