Distribution, Regionalization, and Diversity of the dung beetle genus Phanaeus MacLeay (Coleoptera: Scarabaeidae) using Species Distribution Models Author Lizardo, Viridiana Museo de Zoología, Facultad de Ciencias, Universidad Nacional Autónoma de México, 04510 Mexico City, Mexico. Author Moctezuma, Victor 0000-0002-4532-0302 Centro Tlaxcala de Biología de la Conducta, Universidad Autónoma de Tlaxcala, 90070 La Loma Xicohtencatl, Tlaxcala de Xicohténcatl, Tlaxcala, Mexico. https: // orcid. org / 0000 - 0002 - 4532 - 0302 Author Escobar, Federico 0000-0003-3222-4183 Red de Ecoetología, Instituto de Ecología, A. C., 91063 Xalapa, Veracruz, Mexico. https: // orcid. org / 0000 - 0003 - 3222 - 4183 Corresponding author. lizardo. viridiana @ gmail. com; https: // orcid. org / 0000 - 0003 - 3958 - 3373 lizardo.viridiana@gmail.com text Zootaxa 2022 2022-11-29 5213 5 546 568 journal article 10.11646/zootaxa.5213.5.4 1175-5326 7381694 449781B5-94E0-4B6C-9F6B-D0711FC08BB2

TABLE 1. Phanaeus species checklist, updated species groups and selected variables used in the Species Distribution Modelling

Species	Annual Mean Temperature Annual Precipitation Isothermality Max Temperature of Warmest Month Mean Diurnal Range Mean Temperature of Coldest Quarter Mean Temperature of Warmest Quarter Min Temperature of Coldest Month Precipitation of Driest Month Precipitation of Driest Quarter Precipitation of Wettest Month Precipitation of Wettest Quarter Precipitation Seasonality Temperature Annual Range Temperature Seasonality
P. amethystinus species group
P. amethystinus Harold	X	X			X	X	X		X	X	X
P. blackalleri Delgado-Castillo	X		X	X			X		X		X	X	X
P. genieri Arnaud				X X	X	X X		X		X	X		X
P. guatemalensis Harold			X	X X	X	X		X		X	X		X
P. melampus Harold	X		X	X X			X		X		X	X	X
P. tepanensis Bates*
P. beltianus species group
P. beltianus Bates				X X			X		X		X	X	X
P. sallei Harold	X	X			X	X	X	X	X	X	X	X
P. beltianus Bates		X	X	X X			X		X		X		X
P. changdiazi Kohlmann & Solís*
P. howdeni Arnaud*
P. bispinus species group
P. bispinus Bates	X	X	X	X X			X		X		X	X	X
P. sororibispinus Edmonds & Zídek	X	X	X	X X			X		X				X
P. chalcomelas species group
P. achilles Boheman	X	X		X			X		X		X	X	X
P. cambeforti Arnaud	X	X	X	X			X		X		X	X	X
P. chalcomelas (Perty)		X	X	X X			X		X		X	X	X
P. lecourti Arnaud			X	X X			X		X		X	X
P. meleagris Blanchard			X	X X			X		X		X
P. endymion species group
P. arletteae Arnaud			X	X X	X	X	X	X		X	X		X
P. bravoensis Moctezuma, Sánchez-Huerta & Halffter	X		X	X			X		X		X		X
P. chiapanecus Moctezuma & Halffter					X	X	X		X			X	X
P. edmondsi Moctezuma, Deloya & Halffter			X	X		X	X		X		X		X
P. endymion Harold	X		X	X X			X		X		X	X	X
P. funereus Balthasar			X	X X	X	X		X		X	X		X
P. halffterorum Edmonds		X	X	X		X	X		X				X