Three new goatfishes of the genus Upeneus from the Eastern Indian Ocean and Western Pacific, with an updated taxonomic account for U. itoui (Mullidae: japonicus-species group) Author Uiblein, Franz Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway & National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda, South Africa & Natural History Museum, University of Oslo, Norway & Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, Ha Noi, Vietnam Author Motomura, Hiroyuki The Kagoshima University Museum, Korimoto, Kagoshima, Japan. text Zootaxa 2021 2021-03-01 4938 3 298 324 journal article 7893 10.11646/zootaxa.4938.3.2 640aac64-3963-4435-8e04-7af89edd428b 1175-5326 4569653 DF5541CF-E4BE-43DF-BE55-615F4D4CEFBD

TABLE 1. Species of Upeneus known from the CEITPAS area before this study. Species of the japonicus group are emphasized in bold.

Species	Occurrence in the CEITPAS area	Occurrence elsewhere
Upeneus asymmetricus Lachner, 1954 1	Lombok, S Indonesia; Central Philippines
Upeneus farnis Uiblein & Peristiwady, 2017 2	NE Sulawesi, N Indonesia
Upeneus guttatus (Day, 1868) 3	E India to Japan	WIO; Australia to New Caledonia
Upeneus heemstra Uiblein & Gouws, 2014 4	Chennai (Madras), SE India	WIO
Upeneus heterospinus Uiblein & Pavlov, 2019 5	Lombok, S Indonesia; Central Philippines; South China Sea to S Japan
Upeneus itoui Yamashita, Golani & Motomura, 2011 6, 7	S Japan, Taiwan
Upeneus japonicus (Houttuyn, 1782) 8	South China Sea to Japan and Peter the Great Bay, E Rus- sia; Philippines
Upeneus lombok Uiblein & White, 2015 1	Lombok, S Indonesia
Upeneus luzonius Jordan & Seale, 1907 9	Central and N Philippines
Upeneus margarethae Uiblein & Heemstra, 2010 5	E India to W Thailand	WIO; NW and N Australia to inner Gulf of Carpentaria
Upeneus moluccensis (Bleeker, 1855) 9	E India to S Japan	WIO; Australia to New Caledonia; E Mediterranean a
Upeneus nigromarginatus Bos, 2014 10	S and Central Philippines
Upeneus quadrilineatus Cheng & Wang, 1963 11	Indonesia to S Japan
Upeneus spottocaudalis Uiblein & Gledhill, 2017 2	S Indonesia	NE Australia
Upeneus stenopsis Uiblein & McGrouther, 2012 12	Quezon, Central Philippines	N to NE Australia
Upeneus subvittatus (Temminck & Schlegel, 1843) 12	W Indonesia to S Japan
Upeneus sulphureus Cuvier, 1829 9	E India to S Japan	WIO; Australia to New Caledonia and Fiji b
Upeneus sundaicus (Bleeker, 1855) 9, 13	Myanmar to Hainan, S China; Indonesia; Philippines	Mauritius and Persian Gulf, WIO; Australia to Papua New Guinea
Upeneus supravittatus Uiblein & Heemstra, 2010 14, 15	E India to Myanmar	S Iran to W Sri Lanka, WIO
Upeneus taeniopterus Cuvier, 1829 16	E Sri Lanka; Macao, S China; Ryukyu Islands, S Japan	Pinda Bank, N Mozambique and Rodrigues to Laccadives, WIO; Cocos Islands, SE Indian Ocean; Biak c, E Indonesia; Samoa and Palau to Hawaii, French Polynesia and Cook Islands
Upeneus tragula Richardson, 1846 4	Andaman Islands, E India to Japan	Australia to New Caledonia, Papua New Guinea and Palau
Upeneus vittatus (Forsskål, 1775) 11, 14	E India to Japan	WIO; Australia to Hawaii and Marquesas