Description of a new subterranean breeding brown frog (Ranidae: Rana) from Japan Author Eto, Koshiro Author Matsui, Masafumi 0000-0003-2032-2528 fumi@zoo.zool.kyoto-u.ac.jp Author Sugahara, Takahiro 0000-0002-0858-6339 gahasura@t.vodafone.ne.jp text Zootaxa 2022 2022-11-18 5209 4 401 425 journal article 189448 10.11646/zootaxa.5209.4.1 14cfc8f7-4c7f-4709-8492-a5cdda590eda 1175-5326 7333801 B8AA89CC-5B39-40EB-A385-FF2976DDDE6E

TABLE 1. Comparisons of SVL and the other measurements (means followed by ranges in parenthesis) of two Rana species.

	Rana kyoto sp. n.				R. tagoi
	lineage A-1b-sympatric		A-1b-allopatric		A-1a (R. tagoi topotypes)		A-1a-sympatric		B-2a-sympatric		B-2a-allopatric
	male	female	male	female	male	female	male	female	male	female	male	female
SVL	31.5	31.4	41.3	48.1	43.0	41.8	48.9	48.9	48.1	47.8	44.2	41.2
	(28.8–35.9)	(27.4–36.9)	(36.0–43.9)	(48.0–48.3)	(38.5–45.5)	(40.6–43.0)	(40.2–52.9)	-	(39.5–54.8)	(41.2–54.5) (38.6–52.2)		(33.0–50.2)
N	53	33	7	2	12	7	17	1	31	26	47	25
HL	11.1	11.5	13.5	18.4	15.5	15.4	18.0		17.3		16.3
	(10.2–12.1)	(10.6–13.2)	(12.9–14.4)	(18.4–18.4)	(14.7–16.3)	(14.4–16.1)	(14.4–19.3)		(14.6–20.6)		(13.6–17.8)
HW	11.4	11.0	14.6	18.5	16.6	15.5	18.8		17.7		17.2
	(10.0–12.8)	(9.6–12.6)	(13.6–15.7)	(18.5–18.6)	(14.8–18.5)	(14.8–16.1)	(15.0–20.6)		(14.3–21.2)		(13.4–19.9)
SL	4.1	4.1	4.9	6.2	5.6	5.6	6.8		6.1		6.0
	(3.4–4.7)	(3.6–4.8)	(4.8–5.2)	(6.0–6.5)	(5.2–6.2)	(5.1–5.9)	(5.5–7.5)		(4.9–6.9)		(5.1–6.9)
SNL	2.2	2.1	2.4	3.6	2.9	3.0	3.5		3.3		2.9
	(1.9–2.5)	(1.8–2.5)	(2.2–2.6)	(3.5–3.7)	(2.8–3.1)	(2.9–3.2)	(3.4–3.6)		(2.7–3.8)		(2.2–3.4)
NEL	1.8	1.8	2.1	2.3	2.2	2.3	2.6		2.5		2.5
	(1.6–2.1)	(1.5–2.0)	(2.0–2.3)	(2.3–2.3)	(1.8–2.5)	(2.2–2.4)	(2.6–2.6)		(1.9–3.2)		(1.9–3.1)
EL	4.8	4.8	5.7	7.1	6.1	5.6	6.7		6.6		6.6
	(4.3–5.8)	(4.3–5.2)	(5.5–5.9)	(7.0–7.1)	(5.1–6.9)	(5.2–6.6)	(6.1–7.3)		(5.2–8.1)		(5.3–7.5)
TD	1.4	1.7	1.8	2.7	2.1	2.3	2.7		2.9		1.7
	(0.7–2.0)	(1.1–2.1)	(1.5–2.0)	(2.6–2.8)	(1.3–2.8)	(1.8–2.6)	(2.2–3.7)		(1.9–4.1)		(1.1–2.1)
IND	3.5	3.5	3.9	5.0	4.6	4.4	4.8		4.9		4.7
	(3.1–3.9)	(3.2–3.9)	(3.6–4.2)	(4.9–5.2)	(4.0–5.1)	(4.2–4.6)	(4.2–5.3)		(4.1–5.8)		(3.8–5.3)
IOD	2.3	2.5	2.6	3.3	3.1	3.4	3.6		3.3		3.1
	(1.9–2.9)	(2.0–2.9)	(2.1–2.9)	(3.2–3.3)	(2.5–3.7)	(2.8–3.8)	(3.1–4.0)		(2.5–3.9)		(2.5–3.9)
UEW	3.0	3.0	3.4	4.1	3.9	3.8	4.8		4.4		4.5
	(2.7–3.4)	(2.6–3.5)	(3.3–3.6)	(4.1–4.2)	(3.5–4.4)	(3.4–4.1)	(4.0–5.1)		(3.4–5.1)		(3.7–4.9)
LAL	14.4	13.7	17.0	20.3	19.0	18.0	22.6		21.3		20.9
	(13.2–15.6)	(12.7–15.0)	(16.2–18.5)	(20.3–20.4)	(18.0–20.0)	(16.6–18.9)	(19.0–24.6)		(17.3–24.2)		(16.8–23–1)
HLL	53.9	53.3	62.9	78.7	73.3	72.2	87.1		81.9		81.4
	(48.4–61.0)	(48.7–57.6)	(58.6–67.4)	(78.6–78.8)	(67.1–78.7)	(68.5–74.2)	(71.4–91.6)		(65.3–89.5)		(2–65–88.2)
											......continued on the next page