Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China Author Qi, Shuo 0000-0002-2924-6093 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & qishuo 1992 @ outlook. com; https: // orcid. org / 0000 - 0002 - 2924 - 6093 & These authors contributed equally to this work qishuo1992@outlook.com Author Lyu, Zhi-Tong 0000-0002-4891-3859 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859 & These authors contributed equally to this work lvzht@foxmail.com Author Wang, Jian 0000-0003-4249-7767 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangj- 1994 @ outlook. com; https: // orcid. org / 0000 - 0003 - 4249 - 7767 wangj-1994@outlook.com Author Mo, Yun-Ming Natural History Museum of Guangxi, Nanning, Guangxi 530012, China. moyunming @ 163. com Author Zeng, Zhao-Chi 0000-0003-4054-8399 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399 zhaochizeng@outlook.com Author Zeng, Yang-Jin Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Dai, Ke-Yuan Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Li, Yuan-Qiu Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Grismer, L. Lee 0000-0001-8422-3698 Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California 92515, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698 lgrismer@lasierra.edu Author Wang, Ying-Yong 0000-0003-2024-6641 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangyy @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 2024 - 6641 wangyy@mail.sysu.edu.cn text Zootaxa 2021 2021-12-02 5072 5 401 438 journal article 3143 10.11646/zootaxa.5072.5.1 5758176c-96f6-43b9-b9a9-b2fac894dca8 1175-5326 5748979 B23A6B99-6D47-469F-8A4F-8AD41D77151A

TABLE 1. Literature for morphological characters of 58 recognized species of Boulenophrys.

ID	Species	References
1	B. acuta (Wang, Li & Jin, 2014)	Li et al. 2014; this study
2	B. angka (Wu, Suwannapoom, Poyarkov, Pawangkhanant, Xu, Jin, Murphy & Che, 2019) comb. nov.	Wu et al. 2019
3	B. anlongensis (Li, Lu, Liu & Wang, 2020) comb. nov.	Li et al. 2020b
4	B. baishanzuensis (Wu, Li, Liu, Wang & Wu, 2020) comb. nov.	Wu et al. 2020
5	B. baolongensis (Ye, Fei & Xie, 2007)	Ye et al. 2007; Fei & Ye 2016; Fei 2020
6	B. binchuanensis (Ye & Fei, 1995)	Fei & Ye 2016; Fei 2020
7	B. binlingensis (Jiang, Fei & Ye, 2009) comb. nov.	Fei & Ye 2016; Fei 2020; this study
8	B. boettgeri (Boulenger, 1899)	Fei & Ye 2016; Fei 2020; this study
9	B. brachykolos (Inger & Romer, 1961)	Fei & Ye 2016; Fei 2020; this study
10	B. caobangensis (Nguyen, Pham, Nguyen, Luong & Ziegler, 2020) comb. nov.	Nguyen et al. 2020
11	B. caudoprocta (Shen, 1994) comb. nov.	Fei & Ye 2016; Fei 2020; this study
12	B. congjiangensis (Luo, Wang, Wang, Lu, Wang, Deng & Zhou, 2021) comb. nov.	Luo et al. 2021
13	B. cheni (Wang & Liu, 2014)	Wang et al. 2014; this study
14	B. chishuiensis (Xu, Li, Liu, Wei & Wang, 2020) comb. nov.	Xu et al. 2020
15	B. daiyunensis (Lyu, Wang & Wang, 2021) comb. nov.	Lyu et al. 2021; this study
16	B. daoji (Lyu, Zeng, Wang & Wang, 2021) comb. nov.	Lyu et al. 2021; this study
17	B. daweimontis (Rao & Yang, 1997) comb. nov.	Fei & Ye 2016; Fei 2020
18	B. dongguanensis (Wang & Wang) comb. nov.	Wang et al. 2019a
19	B. fansipanensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018) comb. nov.	Tapley et al. 2018a
20	B. frigida (Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong & Rowley, 2021) comb. nov.	Tapley et al. 2021
21	B. hoanglienensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong & Rowley, 2018) comb. nov.	Tapley et al. 2018a
22	B. huangshanensis (Fei & Ye, 2005)	Fei & Ye 2016; Fei 2020; this study
23	B. insularis (Wang, Liu, Lyu, Zeng & Wang, 2017) comb. nov.	Wang et al. 2017a
24	B. jiangi (Liu, Li, Wei, Xu, Cheng, Wang & Wu, 2020) comb. nov.	Liu et al. 2020
25	B. jingdongensis (Fei & Ye, 1983) comb. nov.	Fei & Ye 2016; Fei 2020; this study
26	B. jinggangensis (Wang, 2012) comb. nov.	Wang et al. 2012; this study
27	B. jiulianensis (Wang, Zeng, Lyu & Wang) comb. nov.	Wang et al. 2019a
28	B. kuatunensis (Pope, 1929)	Fei & Ye 2016; Fei 2020; Tapley et al. 2017; this study
29	B. leishanensis (Li, Xu, Liu, Jiang, Wei & Wang, 2018) comb. nov.	Li et al. 2018
30	B. lushuiensis (Shi, Li, Zhu, Jiang, Jiang & Wang, 2021) comb. nov.	Shi et al. 2021
31	B. liboensis (Zhang, Li, Xiao, Li, Pan, Wang, Zhang & Zhou, 2017) comb. nov.	Zhang et al. 2017
32	B. lini (Wang & Yang, 2014)	Wang et al. 2014; this study
33	B. lishuiensis (Wang, Liu & Jiang, 2017)	Wang et al. 2017b
34	B. minor (Stejneger, 1926)	This study
35	B. mirabilis (Lyu, Wang &Zhao, 2020) comb. nov.	Lyu et al. 2020
36	B. mufumontana (Wang, Lyu & Wang, 2019) comb. nov.	Wang et al. 2019a
37	B. nankunensis (Wang, Zeng & Wang, 2019) comb. nov.	Wang et al. 2019a
38	B. nanlingensis (Lyu, Wang, Liu & Wang, 2019) comb. nov.	Wang et al. 2019a
39	B. obesa (Wang, Li & Zhao, 2014)	Li et al. 2014
40	B. ombrophila (Messenger & Dahn, 2019) comb. nov.	Messenger et al. 2019
41	B. omeimontis (Liu, 1950)	Fei & Ye 2016; Fei 2020
42	B. palpebralespinosa (Bourret, 1937) comb. nov.	Fei & Ye 2016; Fei 2020
43	B. qianbeinsis (Su, Shi, Wu, Li, Yao, Wang & Li, 2020) comb. nov.	Su et al. 2020
44	B. rubrimera (Tapley, Cutajar, Mahony, Chung, Dau, Nguyen, Luong & Rowley, 2017) comb. nov.	Tapley et al. 2017; 2018b
45	B. sangzhiensis (Jiang, Ye & Fei, 2008) comb. nov.	Jiang et al. 2008; Fei & Ye 2016; Fei 2020; this study
46	B. sanmingensis (Lyu & Wang, 2021) comb. nov.	Lyu et al. 2021; this study
47	B. shimentaina (Lyu, Liu, & Wang, 2020) comb. nov.	Lyu et al. 2020; this study
48	B. shuichengensis (Tian & Sun, 1995) comb. nov.	Tian et al. 2000; Fei & Ye 2016; Fei 2020
49	B. shunhuangensis (Wang, Deng, Liu, Wu & Liu, 2019) comb. nov.	Wang et al. 2019b
50	B. spinata (Liu & Hu, 1973)	Fei & Ye 2016; Fei 2020; this study
51	B. tongboensis (Wang & Lyu, 2021) comb. nov.	Lyu et al. 2021; this study
52	B. tuberogranulatus (Shen, Mo & Li, 2010)	Mo et al. 2010; Fei & Ye 2016; Fei 2020; this study
53	B. wugongensis (Wang, Lyu & Wang, 2019) comb. nov.	Wang et al. 2019a
54	B. wuliangshanensis (Ye & Fei, 1995)	Fei & Ye 2016; Fei 2020; this study
55	B. wushanensis (Ye & Fei, 1995)	Fei & Ye 2016; Fei 2020; this study
56	B. xiangnanensis (Lyu, Zeng & Wang, 2020) comb. nov.	Lyu et al. 2020; this study
57	B. xianjuensis (Wang, Wu, Peng, Shi, Lu & Wu, 2020) comb. nov.	Wang et al. 2020
58	B. yangmingensis (Lyu, Zeng & Wang, 2020) comb. nov.	Lyu et al. 2020; this study