Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China Author Qi, Shuo 0000-0002-2924-6093 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & qishuo 1992 @ outlook. com; https: // orcid. org / 0000 - 0002 - 2924 - 6093 & These authors contributed equally to this work qishuo1992@outlook.com Author Lyu, Zhi-Tong 0000-0002-4891-3859 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859 & These authors contributed equally to this work lvzht@foxmail.com Author Wang, Jian 0000-0003-4249-7767 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangj- 1994 @ outlook. com; https: // orcid. org / 0000 - 0003 - 4249 - 7767 wangj-1994@outlook.com Author Mo, Yun-Ming Natural History Museum of Guangxi, Nanning, Guangxi 530012, China. moyunming @ 163. com Author Zeng, Zhao-Chi 0000-0003-4054-8399 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399 zhaochizeng@outlook.com Author Zeng, Yang-Jin Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Dai, Ke-Yuan Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Li, Yuan-Qiu Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Grismer, L. Lee 0000-0001-8422-3698 Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California 92515, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698 lgrismer@lasierra.edu Author Wang, Ying-Yong 0000-0003-2024-6641 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangyy @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 2024 - 6641 wangyy@mail.sysu.edu.cn text Zootaxa 2021 2021-12-02 5072 5 401 438 journal article 3143 10.11646/zootaxa.5072.5.1 5758176c-96f6-43b9-b9a9-b2fac894dca8 1175-5326 5748979 B23A6B99-6D47-469F-8A4F-8AD41D77151A

TABLE 3. Diagnostic characters separating all 61 species of the Boulenophrys.

ID	Species	SVL in	SVL in	Horn-like	Vomerine	Tongue:	Heels:	Lateral fringes	Webs on toes:
		males (in mm)	females (in mm)	tubercle at upper eyelid: slightly	teeth: present (1),	notched (1), or not notched (0)	overlapping (2) just meeting (1),	on toes: wide (2), narrow (1),	more than one- fourth (2),
				large (2), small (1)	or absent (0)		or not meeting (0)	or absent (0)	rudimentary (1), or lacking (0)
1	B. yaoshanensis sp. nov.	32.5–42.6	46.6–47.4	1	0	0	1 or 2	0	1
2	B. yingdeensis sp. nov.	33.2–35.3	36.3–45.8	1	1	0	1 or 2	0	1
3	B. yunkaiensis sp. nov.	35.3–40.0	45.3–46.1	1	0	0	1 or 2	0	1
4	B. acuta	27.1–33.0	28.1–33.6	2	0	0	0	1	1
5	B. angka	31.2–32.1	37.5–39.2	1	0	0	1 or 2	0	1
6	B. anlongensis	40.0–45.5	48.9–51.2	1	0	0	2	1	1
7	B. baishanzuensis	28.4–32.4	/	1	0	0	2	1	0
8	B. baolongensis	42.0–45.0	/	1	0	1	1	0	0
9	B. binchuanensis	32.0–36.0	40.2–42.5	1	0	0 or 1	1	2	1
10	B. binlingensis	45.1–51.0	/	1	0	1	2	/	1
11	B. boettgeri	34.5–37.8	39.7–46.8	1	0	1	1 or 2	2	1
12	B. brachykolos	33.7–39.3	33.9–45.9	1	0	0	0	0	1
13	B. caobangensis	34.9–38.9	/	1	0	0	/	0	1
14	B. caudoprocta	81.3	/	2	1	0	1	/	1
15	B. congjiangensis	28.6–33.4	38.4–40.2	1	0	0	2	1	1
16	B. cheni	26.2–29.5	31.8–34.1	1	0	1	2	2	1
17	B. chishuiensis	43.4–44.1	44.8–49.8	1	0	0	2	0	1
18	B. daiyunensis	27.6–28.7	33.7–35.6	1	1	0	1 or 2	1	1
19	B. daoji	32.6–33.6	37.5–41.4	1	0	0	0	1	1
20	B. daweimontis	34.0–37.0	40.0–46.0	1	1	/	/	0	0
21	B. dongguanensis	30.2–39.3	/	1	1	0	0	0	1
22	B. fansipanensis	30.9–44.3	41.7–42.5	1	1	0 or 1	/	0	0
23	B. frigida	30.3–31.8	/	1	1	0 or 1	/	0	0