Three new species of the genus Boulenophrys (Anura, Megophryidae) from southern China Author Qi, Shuo 0000-0002-2924-6093 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & qishuo 1992 @ outlook. com; https: // orcid. org / 0000 - 0002 - 2924 - 6093 & These authors contributed equally to this work qishuo1992@outlook.com Author Lyu, Zhi-Tong 0000-0002-4891-3859 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & lvzht @ foxmail. com; https: // orcid. org / 0000 - 0002 - 4891 - 3859 & These authors contributed equally to this work lvzht@foxmail.com Author Wang, Jian 0000-0003-4249-7767 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangj- 1994 @ outlook. com; https: // orcid. org / 0000 - 0003 - 4249 - 7767 wangj-1994@outlook.com Author Mo, Yun-Ming Natural History Museum of Guangxi, Nanning, Guangxi 530012, China. moyunming @ 163. com Author Zeng, Zhao-Chi 0000-0003-4054-8399 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & zhaochizeng @ outlook. com; https: // orcid. org / 0000 - 0003 - 4054 - 8399 zhaochizeng@outlook.com Author Zeng, Yang-Jin Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Dai, Ke-Yuan Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Li, Yuan-Qiu Guangdong Shimentai National Nature Reserve, Qingyuan, Guangdong 513000, China Author Grismer, L. Lee 0000-0001-8422-3698 Herpetology Laboratory, Department of Biology, La Sierra University, Riverside, California 92515, USA & Department of Herpetology, San Diego Natural History Museum, PO Box 121390, San Diego, California, 92112, USA & lgrismer @ lasierra. edu; https: // orcid. org / 0000 - 0001 - 8422 - 3698 lgrismer@lasierra.edu Author Wang, Ying-Yong 0000-0003-2024-6641 State Key Laboratory of Biocontrol / The Museum of Biology, School of Ecology / School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong 510275, China & wangyy @ mail. sysu. edu. cn; https: // orcid. org / 0000 - 0003 - 2024 - 6641 wangyy@mail.sysu.edu.cn text Zootaxa 2021 2021-12-02 5072 5 401 438 journal article 3143 10.11646/zootaxa.5072.5.1 5758176c-96f6-43b9-b9a9-b2fac894dca8 1175-5326 5748979 B23A6B99-6D47-469F-8A4F-8AD41D77151A

APPENDIX III. Mean p -distance of 16S gene among the Boulenophrys species used in this study.

ID	Species	1	2	3	4	5	6	7	8	9	10	11	12	13	14	15	16	17	18	19	20
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1	B. yaoshanensis sp. nov.	0.3
2	B. yingdeensis sp. nov.	4.2	0.0
3	B. yunkaiensis sp. nov.	2.8	5.6	0.0
4	B. acuta	6.8	8.1	7.5	/
5	B. angka	6.4	9.0	7.4	10.6	/
6	B. anlongensis	4.7	6.8	6.5	9.1	4.9	/
7	B. baishanzuensis	3.6	5.6	4.1	7.8	7.4	6.2	/
8	B. baolongensis	4.0	5.9	3.8	7.5	7.8	6.6	4.7	/
9	B. binchuanensis	4.6	5.6	5.3	7.8	5.5	3.5	5.9	5.0	/
10	B. binlingensis	5.2	5.6	5.6	8.4	5.8	4.4	5.9	4.4	2.6	/
11	B. boettgeri	2.5	5.2	2.6	6.8	6.5	5.6	2.0	3.2	4.4	4.4	/
12	B. brachykolos	5.5	6.5	6.8	8.8	10.0	6.2	6.5	5.9	5.9	7.8	5.9	/
13	B. caobangensis	6.1	6.5	6.2	9.7	5.2	4.4	6.5	6.8	4.0	4.3	5.6	8.4	/
14	B. caudoprocta	4.0	6.8	3.5	6.9	7.1	6.5	4.4	3.2	5.3	4.7	2.9	7.1	6.8	/
15	B. cheni	3.0	3.5	3.4	8.1	7.1	5.6	3.2	3.2	4.9	5.0	2.9	5.3	5.9	4.3	/
16	B. chishuiensis	5.5	7.4	5.9	9.1	7.4	5.9	5.3	6.2	5.2	5.6	4.7	7.2	6.5	5.6	5.3	/
17	B. congjiangensis	2.8	5.5	3.4	6.5	7.4	6.5	4.6	3.2	4.3	5.8	3.1	5.5	6.5	3.5	4.3	6.2	/
18	B. daiyunensis	4.9	5.9	4.7	8.1	8.7	7.1	4.1	5.6	5.3	5.9	4.1	7.2	7.1	4.4	4.1	5.3	5.5	/
19	B. daoji	3.9	5.2	4.0	8.4	7.4	7.1	5.2	5.0	6.1	6.5	3.7	5.3	6.5	4.9	3.7	6.8	4.9	5.6	/
20	B. daweimontis	4.4	5.2	5.0	6.5	5.2	4.1	4.3	5.0	3.1	3.4	4.0	8.0	4.3	5.3	4.6	6.1	4.9	5.5	5.5	/
21	B. dongguanensis	5.1	5.6	5.9	8.4	9.0	7.8	4.4	5.6	6.5	6.5	5.0	7.8	7.5	6.8	3.2	7.5	7.1	5.9	6.5	5.9
22	B. fansipanensis	4.8	5.8	6.1	10.9	6.1	5.8	5.8	5.0	4.9	4.9	4.9	7.7	5.8	5.8	4.9	5.5	5.2	6.2	5.5	5.5
23	B. frigida	4.5	6.2	4.6	8.4	5.8	5.2	5.5	4.7	4.0	4.6	4.0	7.1	5.2	4.7	4.0	5.5	4.0	5.5	5.5	4.6
24	B. hoanglienensis	4.9	5.3	5.3	8.7	6.4	4.9	5.6	4.7	3.1	2.9	4.1	7.4	4.9	5.0	4.6	4.9	5.5	5.3	6.2	4.0
25	B. huangshanensis	2.9	5.9	3.2	6.8	7.1	5.6	3.2	3.8	5.0	5.0	1.1	6.5	6.2	3.5	3.5	5.3	3.7	4.7	4.3	4.4
26	B. insularis	4.5	4.6	6.1	9.3	8.3	7.4	4.6	5.2	6.7	6.2	4.9	6.8	7.1	6.4	3.1	6.5	6.4	5.0	5.2	6.4
27	B. jiangi	9.9	11.3	10.0	12.4	15.0	13.7	10.9	10.7	11.2	12.6	10.3	12.5	14.0	11.7	10.9	9.8	9.6	10.7	11.3	11.9
28	B. jingdongensis	5.2	6.5	5.9	8.4	5.0	4.4	6.2	5.3	3.8	3.5	5.3	8.8	4.6	5.0	5.3	6.2	5.3	6.8	5.9	3.5
29	B. jinggangensis	4.2	3.8	5.0	6.8	8.0	5.2	5.0	5.0	4.6	4.9	4.1	5.9	5.8	5.6	3.5	5.8	4.9	4.4	4.9	4.9
30	B. jiulianensis	3.6	5.8	2.9	8.7	7.7	6.8	4.3	4.1	5.6	5.0	2.9	7.1	6.2	3.8	4.0	5.9	3.7	4.7	4.9	5.8
31	B. kuatunensis	3.6	4.9	4.4	8.7	7.1	5.9	2.6	4.4	5.6	5.6	2.9	5.3	5.9	4.7	2.9	5.0	4.9	4.4	3.7	5.2
32	B. leishanensis	2.8	5.5	3.2	7.1	6.2	5.3	3.4	2.9	3.8	4.1	2.0	5.9	5.9	3.5	3.1	5.0	2.8	4.4	4.3	4.3
33	B. liboensis	3.6	6.5	3.8	8.1	7.7	5.9	3.4	4.4	5.6	5.6	2.3	5.9	6.2	4.1	4.0	4.7	4.3	5.3	4.3	5.2
34	B. lini	4.8	4.0	5.9	7.4	8.3	6.2	4.0	6.5	6.2	6.8	4.6	5.2	7.1	5.9	4.0	7.1	5.5	5.6	5.5	5.8
35	B. lishuiensis	4.9	5.9	5.3	8.8	8.1	7.2	4.4	4.7	5.6	5.6	3.5	7.2	6.9	5.0	4.4	6.9	5.6	5.6	4.7	5.3
36	B. lushuiensis	6.1	7.8	6.2	9.8	6.2	5.3	6.8	5.6	4.1	4.4	5.3	8.8	5.3	5.3	5.9	6.5	5.9	6.5	7.4	4.4
37	B. minor	7.0	8.8	7.5	9.1	7.7	7.2	6.2	7.9	6.2	7.2	5.6	8.8	7.4	7.2	6.9	3.7	7.8	8.2	8.4	6.5
38	B. mirabilis	3.1	5.6	3.2	7.5	9.0	6.9	4.1	5.0	5.6	6.2	3.5	7.2	7.2	4.4	4.4	6.2	3.8	5.0	5.6	5.6
39	B. mufumontana	4.0	6.2	4.1	8.8	8.1	6.9	4.1	4.7	5.6	5.6	2.6	7.2	6.9	4.7	4.4	5.3	4.6	5.0	5.6	5.9
40	B. nankunensis	4.2	5.9	5.3	9.8	8.8	7.2	4.4	4.4	6.8	6.2	4.7	7.2	8.1	5.6	2.3	7.2	5.5	5.3	5.9	5.9
41	B. nanlingensis	5.6	3.8	7.1	7.7	10.0	7.2	5.3	6.9	6.5	6.5	5.6	6.2	7.5	7.5	4.4	8.4	7.1	6.9	6.2	6.5
42	B. obesa	4.2	5.0	5.2	8.4	8.0	7.1	3.5	5.9	7.1	7.1	4.3	7.4	7.4	6.2	2.6	7.4	5.8	5.9	4.6	4.9
43	B. ombrophila	4.6	6.5	6.5	10.4	8.1	7.2	5.0	5.6	6.8	7.2	5.3	7.5	8.1	6.8	3.5	7.5	6.1	6.2	6.5	6.2
44	B. omeimontis	4.9	5.6	6.2	9.7	4.3	2.9	5.9	5.9	2.9	2.6	5.0	7.8	4.9	5.9	4.4	5.9	6.5	6.5	5.9	3.1
45	B. palpebralespinosa	6.5	7.5	7.8	8.1	6.2	5.9	8.1	6.6	5.0	5.6	6.5	8.8	6.8	6.6	7.1	8.7	6.2	9.1	7.8	5.3
46	B. qianbeiensis	5.4	5.3	5.9	8.3	5.8	3.8	5.6	5.0	2.6	1.4	4.7	7.4	3.2	5.2	4.6	4.9	6.1	5.6	6.8	2.8
47	B. rubrimera	4.9	6.8	5.0	9.7	4.7	3.2	4.7	5.3	3.2	3.5	3.2	7.5	3.8	5.6	4.9	5.0	5.8	5.9	5.9	3.2
48	B. sangzhiensis	4.8	5.2	5.9	8.7	5.8	5.0	5.6	4.7	3.1	1.7	4.6	8.0	4.3	4.9	4.6	5.8	6.1	6.2	6.7	3.4
49	B. sanmingensis	4.2	5.5	5.2	8.7	7.7	6.8	2.9	5.6	7.1	6.8	3.7	7.4	7.4	4.9	3.7	6.8	5.8	5.3	5.8	5.5
50	B. shimentaina	5.1	7.1	5.6	9.8	10.0	8.1	5.3	5.9	6.5	6.5	4.4	8.1	8.3	5.9	4.9	6.9	5.8	5.6	6.8	7.4
51	B. shunhuangensis	2.2	4.7	2.9	6.2	7.4	5.3	2.6	3.5	4.1	4.7	2.0	5.6	5.6	3.5	2.9	4.7	2.9	4.1	4.6	4.7
52	B. spinata	5.8	5.3	6.2	9.0	6.8	5.3	5.9	5.0	3.5	1.4	5.0	8.4	4.7	5.2	5.0	6.2	6.4	6.5	7.1	3.7
53	B. tongboensis	5.1	5.9	4.4	9.1	8.8	7.8	4.7	4.7	5.6	5.9	3.8	7.5	7.2	5.0	4.1	7.5	5.3	5.3	5.0	5.6
54	B. tuberogranulatus	2.5	4.6	2.9	6.8	6.5	5.0	3.2	2.0	3.8	3.2	1.7	5.3	5.6	2.6	2.3	4.7	3.1	4.1	3.7	4.0
55	B. wugongensis	5.6	5.3	5.0	9.4	8.0	6.8	5.3	5.6	5.3	5.9	5.0	8.1	5.5	5.9	3.2	6.2	6.4	4.7	5.6	4.9
56	B. wuliangshanensis	4.0	5.9	4.7	8.8	5.9	4.4	4.9	5.6	3.8	4.1	4.7	7.8	5.6	5.9	5.2	6.2	4.9	6.5	6.2	3.5
57	B. wushanensis	3.6	4.9	4.3	8.4	7.4	5.9	4.0	3.2	4.3	4.7	3.2	6.2	7.1	4.7	3.7	6.2	4.0	5.6	4.9	4.3
58	B. xiangnanensis	3.9	2.0	5.3	7.8	8.7	5.9	5.3	5.9	5.3	5.9	4.9	5.3	6.8	6.5	3.2	6.8	5.2	5.6	4.6	4.9
59	B. xianjuensis	3.7	5.9	3.8	8.7	6.5	6.2	4.3	4.7	5.3	5.6	2.9	6.8	5.9	4.7	4.0	7.1	4.6	5.9	2.6	4.6
60	B. yangmingensis	2.9	5.9	3.5	7.1	7.7	5.9	3.8	3.2	5.0	4.4	2.3	6.5	6.8	3.2	3.4	5.3	3.7	4.7	4.9	4.7