The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus Author Paul, Gregory S. 3100 St. Paul St. 604, Baltimore, MD 21218, USA GSP1954@aol.com Author Iv, W. Scott Persons Department of Geology and Environmental Geosciences, College of Charleston, 202 Calhoun St, Charleston, SC 29401, USA Author Van Raalte, Jay 3100 St. Paul St. 604, Baltimore, MD 21218, USA text Evolutionary Biology 2022 2022-03-01 49 2 156 179 https://doi.org/10.1007/s11692-022-09561-5 journal article 20425 10.1007/s11692-022-09561-5 df53e152-ad9e-4b59-b7a7-1ffec806ac46 6323344

Table 2 Measurementsinmmandratiosfor Tyrannosaurus humeri, ilia, metatarsals 2 and 4, maxillae, dentaries, anddentaryteeth 2 and 3, withadditionalcalculations

Tyrannosaurus Hum Leng		Hum Circ	Hum Ratio	Ilium Leng	Ilium Depth	Ilium Ratio Meta2 Leng Meta2 Circ			Meta2 L/C Ratio	Meta2 Diam Meta2 L/D Ratio
BHI 3033				1540	585	2.63	595	280	2.13	69	8.6
Z-rex							610	305	2
RSM P2523.8				1545	515	3
FMNH PR2081	390	185	2.11	1525	608	2.51	584			78	7.49
BHI 6248
TMT v222
MOR 1128
USNM 555000 375		162	2.31	1470	513	2.86	585	295	1.98
CM 9380	330	145	2.28	1515	~530	~ 2.85	615			89	6.91
MOR 980	362	165	2.19	1397	483	2.89	597	232	2.57
RMDRC 2002
HMN MB.R.91216
TMP 81.6.1	302	150	2.01
LL 12823
BHI 6242
LACM 150167
BHI 6232
BHI 6435
BHI 6436
RGM 792.000
MOR 1125
BHI 6233
BHI 6230	330	145	2.28				600	272	2.21	64	9.38
MOR 009				1180	407	2.9	540			70	7.71
USNM 6183
LACM 23845							465			48	9.69
LACM 23844							575			75	7.67
BHI 4100
BHI 4182
BHI 6231	360	172	2.09
MOR 008
AMNH 5027				1448	470	3.08
NHMUK R7994
SDSM 12047