New genus and species of Philippine spittlebug (Cercopidae: Hemiptera), with notes on subtribal placement of Trigonoschema Author Jr Crispolon, Elorde S. Department of Crop Protection, Division of Entomology, College of Agriculture, University of Southern Mindanao, Kabacan, Cotabato, Philippines. & Mécanismes adaptatifs et évolution (MECADEV), Muséum national d’Histoire naturelle, CNRS, CP 50, Entomologie, 57 rue Cuvier, 75005 Paris, France. ejscrispolon@usm.edu.ph Author Yap, Sheryl A. Institute of Weed Science, Entomology and Plant Pathology, College of Agriculture and Food Science and Museum of Natural History, University of the Philippines Los Baños, Laguna, Philippines. sayap3@up.edu.ph Author Soulier-Perkins, Adeline Mécanismes adaptatifs et évolution (MECADEV), Muséum national d’Histoire naturelle, CNRS, CP 50, Entomologie, 57 rue Cuvier, 75005 Paris, France. adeline.soulier@mnhn.fr text European Journal of Taxonomy 2026 2026-04-28 1052 82 104 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3270/14443 journal article 315114 10.5852/ejt.2026.1052.3270 854e6754-8f2b-4e52-9047-bdde4ae62d1d 2118-9773 19955526 1FDDE331-0EC8-4E37-9BD3-58BE5AC8BE02
Table 3. Character diagnosis for the three closely related genera from the Philippines (modified from Crispolon et al. 2021).
Genera
No.CharactersMioscartaPoeciloterpaTrigonoschemaMiosterpa Crispolon & Yap gen. nov.
1habitus general shape in lateral viewdorso-ventrally flattened, total length nearly 4 × heightglobulous, total length around 2.5 × heightnot dorso-ventrally flattened, in lateral view total length around 3 × heightdorso-ventrally flattened, in lateral view total length around 3.5 × height (Figs 1, 2A)
2total length of the specimen8–11 mmnot exceeding 8 mm9.5–12.5 mm6–7 mm
3Pronotum anglenot more than 25°42°around 45°around 45°
4distance between ocellus and compound eye2 × ocellus diameter6 × ocellus diameterless than 4 × ocellus diameter3 × ocellus diameter
5distance between ocelli1.5 ocellus diameterat least 2 × ocellus diameter1.5 ocellus diameter1 ocellus diameter
6size of ocelli evaluated according to the distance between eyes using ocellus diameterlarge, distance between eyes less than 8 × ocellus diametervery small, distance between eyes 16 × ocellus diametersmall, distance between eyes 9–10.5 × ocellus diametersmall, distance between eyes 9–10.5 × ocellus diameter
7postclypeus shape in frontal viewovoidcompressed laterally with lateral sides almost straightovoidovoid (Figs 1, 2B)
8lateral carinae on postclypeus in frontal viewabsentpresent, prominentabsentpresent prominent (Figs 1, 2B)
9widest part of postclypeus in frontal viewmid heightclose to fronsclose to fronsmid height
10Apical reticulation of the tegmengenerally developed and reduced in few casesreducedreducedreduced (Figs 1, 2C)
11apical curve of tegmen in dorsal viewvisiblerarely visiblerarely visiblevisible (Figs 1, 2C)
12widest part of habitusmidlength of tegmenbefore midlength of tegmenbefore midlength of tegmenbefore midlength of tegmen (Figs 1, 2C)
13apical cells of the tegmennot concaveconcavenot concaveconcave (Figs 1, 2C)
14Rp posterior wingRp separating from SC+Ra nearly at midlength (Fig. 3A)Rp separating from SC+Ra after midlength and making a strong bend after separating from SC+ Ra (Fig. 3C)Rp separating from SC+Ra nearly at midlength (Fig. 3D)Rp separating from SC+Ra nearly at midlength, (Fig. 3A– B)
15subgenital plates lengthat least 1.5 × as long as pygofer heightslightly longer than pygofer heightclearly shorter than 1.5 × as long as pygofer heightas long as the pygofer height (Figs 1, 2E–F)
16subgenital plates fine appendagealways present, longer than the main plategenerally absent, if present shorter than the main plategenerally present but shorter than the main plateabsent (Figs 1, 2E–F)
17paramere general shapenot globose and without protrusion on lateral sidenot globose and without protrusion on lateral sideglobose with protrusion on lateral sidenot globose and without protrusion on lateral side (Figs 1, 2G)