Taxonomy and trans-Beringian biogeography of the pond snails (Gastropoda: Lymnaeidae) of East Asia: an integrative view Author Aksenova, Olga V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia aksyonova.olga@gmail.com Author Vinarski, Maxim V. Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia Author Itagaki, Tadashi Laboratory of Veterinary Parasitology, Faculty of Agriculture, Iwate University, Morioka, Iwate, Japan Author Ohari, Yuma Laboratory of Veterinary Parasitology, Faculty of Agriculture, Iwate University, Morioka, Iwate, Japan & Laboratory of Parasitology and Zoology, School of Veterinary Medicine, Rakuno Gakuen University, Ebetsu, Japan Author Oshida, Tatsuo Laboratory of Wildlife Biology, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan Author Kim, Sang Ki Nakdonggang National Institute of Biological Resources, Gyeongsangbuk-do, Sangju, 37242, Republic of Korea Author Lee, Jin Hee Daegu Science High School, Daegu, 42110, Republic of Korea Author Kondakov, Alexander V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Khrebtova, Irina S. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Soboleva, Alena A. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Travina, Oksana V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Sokolova, Svetlana E. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Palatov, Dmitry M. Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia & A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, 33 Leninsky Prospekt, 119071 Moscow, Russia Author Bespalaya, Yulia V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia & Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia Author Vikhrev, Ilya V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Gofarov, Mikhail Yu. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Bolotov, Ivan N. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia text Zoological Journal of the Linnean Society 2024 2024-08-03 201 4 1 24 http://dx.doi.org/10.1093/zoolinnean/zlae083 journal article 303639 10.1093/zoolinnean/zlae083 96f31273-7b71-4746-8672-022ce216ed8f 0024-4082 13835601 B848A01-DC8F-4759-91E9-237E4526462C

Table 4. Conchometric characteristics of type specimens of the newly described lymnaeid species and a sample of Orientogalba ollula.

Character /index	Species (number of measured shells)
In each cell, above line,limits of variation (minimum–maximum); below line,mean value ± σ.
	Galba pacifica (N = 14)	Kamtschaticana nipponica	Orientogalba hokkaidoensis	Orientogalba ollula
		(N = 5)	(N = 9)	(N = 10)
Number of whorls	4.25–5.25	3.50–4.00	4.25–5.25	3.37–5.12
	4.72 ± 0.38	3.81 ± 0.24	4.90 ± 0.40	4.19 ± 0.45
Shell height (SH), mm	6.0–8.8	9.3–12.0	9.3–11.7	6.53–9.52
	7.5 ± 1.0	10.0 ± 1.2	10.9 ± 0.7	7.50 ± 0.96
Shell width (SW), mm	3.5–5.2	6.5–8.4	6.1–7.3	4.07–5.69
	3.6 ± 0.8	7.2 ± 0.8	6.7 ± 0.4	4.62 ± 0.56
Spire height (SpH), mm	2.3–4.7	2.2–3.9	2.2–4.2	2.18–3.76
	3.6 ± 0.8	3.0 ± 0.7	3.6 ± 0.7	2.76 ± 0.48
Body whorl height (BWH), mm	4.7–6.5	8.2–10.8	7.1–9.6	4.59–7.95
	5.7 ± 0.6	9.0 ± 1.0	8.7 ± 0.8	5.89 ± 0.85
Aperture height (AH), mm	3.3–4.6	6.5–8.2	5.9–7.8	4.23–7.52
	3.9 ± 0.4	7.1 ± 0.7	7.1 ± 0.6	4.89 ± 1.02
Aperture width (AW), mm	2.1–3.3	6.5–8.2	4.7–5.6	2.56–3.96
	2.7 ± 0.3	5.0 ± 0.7	5.2 ± 0.3	3.24 ± 0.42
Index SW/SH	0.53–0.75	0.69–0.83	0.58–0.66	0.59–0.66
	0.60 ± 0.06	0.72 ± 0.06	0.62 ± 0.03	0.62 ± 0.02
Index SpH/SH	0.38–0.54	0.25–0.34	0.19–0.38	0.33–0.41
	0.48 ± 0.04	0.29 ± 0.05	0.33 ± 0.06	0.37 ± 0.03
Index BWH/SH	0.72–0.83	0.88–0.95	0.62–0.87	0.62–0.84
	0.77 ± 0.03	0.90 ± 0.02	0.80 ± 0.07	0.79 ± 0.08
Index AH/SH	0.45–0.63	0.65–0.76	0.62–0.73	0.59–0.85
	0.52 ± 0.05	0.71 ± 0.05	0.65 ± 0.03	0.65 ± 0.07
Index AW/AH	0.61–0.85	0.62–0.78	0.66–0.83	0.49–0.75
	0.70 ± 0.06	0.70 ± 0.07	0.73 ± 0.04	0.67 ± 0.08