Taxonomy and trans-Beringian biogeography of the pond snails (Gastropoda: Lymnaeidae) of East Asia: an integrative view Author Aksenova, Olga V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia aksyonova.olga@gmail.com Author Vinarski, Maxim V. Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia Author Itagaki, Tadashi Laboratory of Veterinary Parasitology, Faculty of Agriculture, Iwate University, Morioka, Iwate, Japan Author Ohari, Yuma Laboratory of Veterinary Parasitology, Faculty of Agriculture, Iwate University, Morioka, Iwate, Japan & Laboratory of Parasitology and Zoology, School of Veterinary Medicine, Rakuno Gakuen University, Ebetsu, Japan Author Oshida, Tatsuo Laboratory of Wildlife Biology, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan Author Kim, Sang Ki Nakdonggang National Institute of Biological Resources, Gyeongsangbuk-do, Sangju, 37242, Republic of Korea Author Lee, Jin Hee Daegu Science High School, Daegu, 42110, Republic of Korea Author Kondakov, Alexander V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Khrebtova, Irina S. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Soboleva, Alena A. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Travina, Oksana V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Sokolova, Svetlana E. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Palatov, Dmitry M. Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia & A. N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, 33 Leninsky Prospekt, 119071 Moscow, Russia Author Bespalaya, Yulia V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia & Laboratory of Macroecology and Biogeography of Invertebrates, Saint Petersburg State University, 7 / 9 Universitetskaya Embankment, 199034 Saint Petersburg, Russia Author Vikhrev, Ilya V. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Gofarov, Mikhail Yu. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia Author Bolotov, Ivan N. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia text Zoological Journal of the Linnean Society 2024 2024-08-03 201 4 1 24 http://dx.doi.org/10.1093/zoolinnean/zlae083 journal article 303639 10.1093/zoolinnean/zlae083 96f31273-7b71-4746-8672-022ce216ed8f 0024-4082 13835601 B848A01-DC8F-4759-91E9-237E4526462C

Table 1. The type series of the new lymnaeid species described in this paper.

Species	N	Sampling site and date	Depository, accession number
Galba pacifica (the holotype and a paratype)	2	Japan, Hokkaido Prefecture, Biei Town, the Ishikari River system, a ditch near the Rubeshibe Stream, 43.520331°N, 142.372452°E, 7 September 2015, leg. Y. Ohari	ZIN, no. 1/531-2021 (holotype), 3/531-2021 (paratype)
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Biei Town, the Ishikari River system, a ditch near the Kawamukai Stream, 43.528123°N, 142.594829°E, 7 September 2015, leg. Y. Ohari	ZIN, no. 2/531-2021
G. pacifica (paratypes)	3	Japan, Hokkaido Prefecture, Hidaka Town, the Saru River system, a ditch near the Pankeusyappu Stream, 42.927325°N, 142.402711°E, 9 September 2015, leg. Y. Ohari	ZIN, no. 3/531-2021
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Shimukappu village, the Mukawu River system, a ditch near the Horokatomamu Stream, 43.058562°N, 142.529372°E, 8 September 2015, leg. Y. Ohari	ZIN, no. 4/531-2021
G. pacifica (paratypes)	1	Japan, Hokkaido Prefecture, Shin-hidaka Town, the Shizunai River system, a ditch near the Sunbetsu Stream, 42.445675°N, 142. 549053°E, 17 September 2013, leg. Y. Ohari	RMBH, no. MLym-1113/7
G. pacifica (paratypes)	1	Japan, Hokkaido Prefecture, Shin-hidaka Town, the Shizunai River system, a ditch near the Sunbetsu Stream, 42.445675°N, 142.549053°E, 17 September 2015, leg. Y. Ohari	RMBH, no. MLym-1113/6
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Nakagawa Town, the Teshio River system, a ditch near the Chirashinai Stream, 44.72604°N, 142.062333°E, 8 August 2015, leg. Y. Ohari	RMBH, no. MLym-1113/2
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Niseko Town, the Shiribetsu River system, a ditch near the Konbu Stream, 42.772299°N, 140.598945°E, 25 May 2015, leg. Y. Ohari	RMBH, no. MLym-1113/1
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Engaru Town, the Yubetsu River system, a ditch near the Shiyubetsu Stream, 43.862232°N, 143.189478°E, 6 September 2015, leg. Y. Ohari	RMBH, no. MLym-1113/5
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Naie Town, the Ishikari River system, Naie Stream, 43.391666°N, 141.975526°E, 3 September 2015, leg. Y. Ohari	RMBH, no. MLym-1113/4
G. pacifica (a paratype)	1	Japan, Hokkaido Prefecture, Wakkanai City, the Koetoi River system, Uruya Stream, 45.261444°N, 141.906737°E, 9 August 2015, leg. Y. Ohari	RMBH, no. MLym-1113/3
Kamtschaticana nipponica (the holotype)	1	Japan, Hokkaido Prefecture, Sarabetsu village, ZIN, no. 1/528-2021 the Tokachi River system, Itarataraki Stream, 42.627161°N, 143.265175°E, 3 October 2013, leg. Y. Ohari
K. nipponica (paratypes)	4	Japan, Hokkaido Prefecture, Sarabetsu village, the ZIN, no. 2/528-2021 Tokachi River system, Saccharobetsu Stream, 42.627161°N, 143.265175°E, 22 June 2013, leg. Y. Ohari
K. nipponica (paratypes)	3	Japan, Hokkaido Prefecture, Obihiro City, the Tokachi River system, Tobetsu Stream, 42.766686°N, 143.202864°E, 18 September 2013, leg. Y. Ohari	RMBH, no. MLym-1115/2, MLym-1115/3, and MLym-1115/4
K. nipponica (a paratype)	1	Japan, Hokkaido Prefecture, Rankoshi Town, a ditch near the Shiribetsu River, 42.815818°N, 140.535244°E, 25 May 2015, leg. Y. Ohari	RMBH, no. MLym-1115/1
Orientogalba hokkaidoensis (the holotype + paratypes)	3	Japan, Hokkaido Prefecture, Hokuto City, a ZIN, no. 1/530-2021 (holotype), ditch near the Hikirichi Stream, 41.842808°N, 2/530-2021 (paratypes) 140.634441°E, 29 June 2015, leg. Y. Ohari
O. hokkaidoensis (a paratype)	1	Japan, Hokkaido Prefecture, Horokanai Town, the Ishikari River system, a ditch near the Uryu Stream, 44.052343°N, 142.142657°E, 4 September 2015, leg. Y. Ohari	ZIN, no. 3/530-2021
O. hokkaidoensis (paratypes)	3	Japan, Hokkaido Prefecture, Tomomae Town, the Kotanbetsu River system, a ditch near the Sankebetsu River, 44.192775°N, 141.771754°E, 7 August 2015, leg. Y. Ohari	RMBH, no. MLym-1114/1, no. MLym-1114/2, and no. MLym-1114/3
O. hokkaidoensis (a paratype)	1	Japan, Hokkaido Prefecture, Kikonai Town, the RMBH, no. MLym-1114/4 Kikonai River system, a ditch near the Uriya Stream, 41.687164°N, 140.386156°E, 29 June 2015, leg. Y. Ohari
O. hokkaidoensis (a paratype)	1	Japan, Hokkaido Prefecture, Fukushima Town, a ditch near the Fukushima River, 41.490724°N, 140.250207°E, 29 June 2015, leg. Y. Ohari	RMBH, no. MLym-1114/5