On the occurrence of the genus Schellencandona (Ostracoda, Candonidae) in Italy, with the description of a new species Author Rossetti, Giampaolo Department of Chemistry, Life Science and Environmental Sustainability, University of Parma, 43124 Parma, Italy Author Mazzini, Ilaria CNR-IGAG, Area della Ricerca di Roma 1, 00015 Montelibretti (RM), Italy Author Bruno, Maria Cristina Research and Innovation Centre, Fondazione Edmund Mach, 38098 S. Michele all’Adige (TN), Italy & NBFC, National Biodiversity Future Center, Palermo 90133, Italy text Zootaxa 2025 2025-11-25 5723 2 189 208 https://doi.org/10.11646/zootaxa.5723.2.2 journal article 314027 10.11646/zootaxa.5723.2.2 7030501b-8a1b-4243-a609-10046b9e8bb9 1175-5326 17893105 7E331561-4D0D-4757-A49F-E3F9F70A3BC7

TABLE 1. Literature-based occurrences of Schellencandona species. The current taxonomic classification follows Meisch et al. (2024). a: reported as Schellencandona dui in Table 2 and as Schellencandona cf. belgica in Plate 3, g; b: see Issartel and Marmonier, 2025a; c: reported as Shellencandona; d: this record, anterior to the description of S. rhodanensis Issartel and Marmonier, 2025a, can most plausibly be assigned to this species.

References	Original taxonomic designation	Current taxonomic classification	Country	Habitat	Recent/fossil	Illustrated material
in Table 2 and as Schellencandona cf. belgica in Plate 3, g; b: see Issartel and Marmonier, 2025a; c: reported as Shellencandona; d: this record, anterior to the description of S. rhodanensis Issartel and Marmonier, 2025a, can most plausibly be assigned to this species.
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Pieri et al. 2009	Schellencandona sp.	Marmocandona sp.	Italy	Alkaline peat bog fed by resurgence springs	Recent	Valves
Petkovski et al. 2009	Schellencandona cf. aemonae	Pseudocandona cf. aemonae	Bosnia and Herzegovina	Cave	Recent	Valves
Issartel & Marmonier 2025b	Schellecandona danielopoli	Schellecandona danielopoli	France	Interstitial	Recent	Valves and soft parts
Marmonier & Creuzé des Châtelliers 1992	Pseudocandona cf. schellenbergi	Schellencandona cf. schellenbergi	France	Interstitial	Recent	No
Ferreira et al. 2003	Schellencandona belgica	Schellencandona belgica	France	Subterranean	Recent	No
Fuhrmann 2012	Schellencandona belgica	Schellencandona belgica	Germany	Cave	Late Holocene	Valves
Fuhrmann & Völker 2015	Schellencandona belgica	Schellencandona belgica	Germany	Cave sediments	Recent	No
Gerecke et al. 2005	Schellencandona belgica	Schellencandona belgica	Luxembourg	Interstitial	Recent	No
Hubart & Dethier 1999	Schellencandona belgica	Schellencandona belgica	Belgium	Springs	Recent	No
Klie 1937	Candona belgica	Schellencandona belgica	Belgium	Wells? Interstitial?	Recent	Valves and soft parts
Marmonier & Creuzé des Châtelliers 1992	Pseudocandona belgica	Schellencandona belgica	France	Interstitial	Recent	No
Marmonier & Creuzé des Châtelliers 1992	Pseudocandona belgica	Schellencandona belgica	Germany	Well	Recent	No
Meisch 2000	Schellencandona belgica	Schellencandona belgica	Germany	Groundwater pipes	Recent	Valves and soft parts
Szlauer-Łukaszewsk 2014	Schellencandona belgica	Schellencandona belgica	Poland	Groyne fields	Recent	No
Szlauer-Łukaszewsk et al. 2021	Schellencandona belgica	Schellencandona belgica	Poland	Springs	Recent	No
Issartel & Marmonier 2025b	Schellencandona capderreyae	Schellencandona capderreyae	France	Interstitial	Recent	Valves and soft parts
Chen et al. 2019	Schellencandona cf. belgica	Schellencandona cf. belgica	China	Lake	Recent	Valves
Sari & Külköylüoğlu 2010	Schellencandona cf. belgica	Schellencandona cf. belgica	Turkey	Unknown	Recent	Valves
Wang et al. 2022	Schellencandona cf. belgica a	Schellencandona cf. belgica	China	Brackish water biofaces	Late Pleistocene	Valves
Ferreira et al. 2003	Schellencandona cf. schellenbergi sp. 1	Schellencandona cf. schellenbergi sp. 1	France	Groundwaters	Recent	No
Ferreira et al. 2003	Schellencandona cf. schellenbergi sp. 2	Schellencandona cf. schellenbergi sp. 2	France	Groundwaters	Recent	No
Dole-Olivier et al. 2006, Prié et al. 2024	Schellencandona cf. simililampadis	Schellencandona cf. simililampadis	France	Springs	Recent	No
Juberthie et al. 2016	Schellencandona cf. tea	Schellencandona cf. tea	Russia	Groundwaters	Recent	No
Issartel & Marmonier 2025b	Schellencandona claretae	Schellencandona claretae	France	Interstitial	Recent	Valves and soft parts
Ma & Yu 2018	Schellencandona dui	Schellencandona dui	China	Ditches with mud, streams with mud and vegetation, lake with vegetation	Recent	Valves and soft parts
Fuchs et al. 2006, Hahn & Fuchs 2009	Schellencandona insueta	Schellencandona insueta	Germany	Boreholes	Recent	No
Klie 1938	Candona insueta	Schellencandona insueta	Germany	Well	Recent	Valves and soft parts (T3 and CR)
Löffler 1961	Candona insueta	Schellencandona insueta	Germany	Interstitial	Recent	No
Yavuzatmaca et al. 2015	Schellencandona insueta	Schellencandona insueta	Turkey	Spring	Recent	No
Issartel & Marmonier 2025b	Schellencandona malardi	Schellencandona malardi	France	Interstitial	Recent	Valves and soft parts
Issartel & Marmonier 2025b	Schellencandona mercantourensis	Schellencandona mercantourensis	France	Interstitial	Recent	Valves and soft parts
Sywula 1976	Candona (Typhlocypris) mira	Schellencandona mira	Poland	Wells	Recent	Valves and some soft parts
Dole-Olivier et al. 2009	Schellencandona triquetra	Schellencandona rhodanensis b	France	Medium and coarse alluvia	Recent	Valves
Issartel & Marmonier 2025a	Schellencandona rhodanensis	Schellencandona rhodanensis	France	Interstitial in main river channels and associated wetlands	Recent	Valves and soft parts
Marmonier & Creuzé des Châtelliers 1992	Pseudocandona triquetra	Schellencandona triquetra b	France	Interstitial	Recent	No
Marmonier et al. 2019	Schellencandona triquetra	Schellencandona rhodanensis b	France	Interstitial	Recent	No
Marmonier et al. 2020	Schellencandona triquetra	Schellencandona rhodanensis b	France	Interstitial and karst cavities	Recent	No
Castellarini et al. 2007	Schellencandona schellenbergi	Schellencandona schellenbergi	France	Streambed sediments?	Recent	No
Fuchs et al. 2006, Fuchs 2007, Hahn & Fuchs 2009	Schellencandona schellenbergi	Schellencandona schellenbergi	Germany	Boreholes	Recent	No
Gibert et al. 1981	Candona aff. schellenbergi	Schellencandona schellenbergi	France	Interstitial	Recent	No
Klie 1934	Candona schellenbergi	Schellencandona schellenbergi	Austria	Wells	Recent	Valves and soft parts
Meisch 1996	Schellencandona schellenbergi	Schellencandona schellenbergi	Germany	Interstitial	Recent	No
Vervier et al. 1986	Pseudocandona schellenbergi	Schellencandona schellenbergi	France	Cave, artificial substrates	Recent	No
Danielopol 1978	Schellencandona similampadis	Schellencandona simililampadis	France	Artificial tunnel in karstic system	Recent	Valves and soft parts
Ferreira et al. 2003	Schellencandona simililampadis	Schellencandona simililampadis	France	Subterranean	Recent	No
Kenderov & Apostolov 2008	Schellencandona sp.	Schellencandona sp.	Bulgaria	Interstitial	Recent	No
Mazzini et al. 2016	Schellencandona sp.	Schellencandona sp.	Albania	Lake	Late Pleistocene	No
Melis et al. 2012	Schellencandona sp.	Schellencandona sp.	Italy	Interstitial?	Holocene	No
Valavani et al. 2024	Schellencandona sp.	Schellencandona sp.	Greece	Cave	Recent	No
Yavuzatmaca et al. 2024	Schellencandona sp.	Schellencandona sp.	Turkey	Stream surface sediment	Recent	No
Dole-Olivier et al. 2009	Schellencandona sp. 4	Schellencandona sp. 4	France	Subterranean	Recent	No
Dole-Olivier et al. 2009	Schellencandona sp. J 1 schellenbergi	Schellencandona sp. J 1 schellenbergi	France	Limestone and coarse alluvium formations	Recent	No
Dole-Olivier et al. 2009	Schellencandona sp. J 2 insueta	Schellencandona sp. J 2 insueta	France	Subterranean	Recent	No
Dole-Olivier et al. 2009	Schellencandona sp. J 3	Schellencandona sp. J 3	France	Subterranean	Recent	No
Khand 2007	Schellencandona sp.?	Schellencandona sp.?	Mongolia	Springs and rivers	Recent	Valves and soft parts
Karanovic & Lee 2012	Schellencandona tea	Schellencandona tea	South Korea	Interstitial	Recent	Valves and soft parts
Dehédin 2012	Schellencandona triquetra c	Schellencandona triquetra	France	Interstitial?	Recent	No
Ferreira et al. 2003	Schellencandona triquetra	Schellencandona triquetra	France	Subterranean	Recent	No
Fuchs et al. 2006, Fuchs 2007, Hahn & Fuchs 2009	Schellencandona triquetra	Schellencandona triquetra	Germany	Boreholes	Recent	No
Fuhrman R. 2012	Schellencandona belgica	Schellencandona belgica	Germany	Boreholes	Late Holocene	Valves
Gerecke et al. 2005	Schellencandona triquetra	Schellencandona triquetra	Luxembourg	Interstitial	Recent	No
Hartmann & Hiller 1977	Candona triquetra	Schellencandona triquetra	Germany	Interstitial	Recent	No
Hubart & Dethier 1999	Schellencandona triquetra	Schellencandona triquetra	Belgium	Springs	Recent	No
Husmann 1956	Candona triquetra	Schellencandona triquetra	Germany	Interstitial	Recent	No
Husmann 1957	Candona triquetra	Schellencandona triquetra	Germany	Intestitial	Recent	No
Husmann 1964	Candona triquetra	Schellencandona triquetra	Germany	Interstitial	Recent	No
Klie 1936	Candona triquetra	Schellencandona triquetra	Belgium	Wells	Recent	Valves and soft parts
Leruth 1937	Candona triquetra	Schellencandona triquetra	Belgium	Cave lake	Recent	No
Muller et al. 2008	Schellencandona triquetra	Schellencandona triquetra d	France	Interstitial	Holocene	Valves
Smith & Kamiya 2006	Schellencandona yakushimaensis	Schellencandona yakushimaensis	Japan	Cracks in bedrock filled with water and sediment	Recent	Valves and soft parts
Smith & Chang 2022	Schellencandona yakushimaensis	Schellencandona yakushimaensis	Japan	Spring	Recent	Valves
Knight 2013, Knight 2017	Mixtacandona / Schellencandona sp.	Unknown	United Kingdom	Cave pool	Recent	No