Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC

Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold.

	D (in mm)	H (in mm)	whorls	D/H	U/D	shell periphery	shell coloration	scale sculpture	spiral lines on apical side	penis	penis coecum	flagellum	node at flagellum
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.
Table 4 (continued). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java.
L. winteriana	9.7–12.6 (mean 11.8)	5.5–7.8 (mean 6.7)	5–5.75 (mean 5.3)	1.55–2.07 (mean 1.76)	0.29–0.51 (mean 0.35)	keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	with large globular bulge at proximal end	absent	distal part crenated, proximal part short	absent
L. conoidea	7.6–8.7 (mean 8.1)	5.2–5.6 (mean 5.3)	4.75–5 (mean 4.9)	1.48–1.54 (mean 1.52)	0.30–0.33 (mean 0.31)	sharply keeled	brownish- corneous	tubercles all over teleoconch	not discernible	?	?	?	?
L. rotatoria	15.6–19.1 (mean 17.4)	8.3–10.6 (mean 9.5)	5.75–6.5 (mean 6.1)	1.57–2.09 (mean 1.85)	0.24–0.32 (mean 0.29)	keeled	brown	scaly processes sparse, at periphery	indistinct	with small globular bulge at proximal end	absent	longish lemon-shaped	absent
L. naggsi sp. nov.	11.6–14.0 (mean 12.5)	8.5–10.6 (mean 9.6)	6.25–7 (mean 6.6)	1.21–1.37 (mean 1.31)	0.28–0.33 (mean 0.31)	angular	brownish- corneous	scaly processes all over teleoconch	not discernible	?	?	?	?
L. epiplatia	11.8–17.0 (mean 15.3)	5.7–6.9 (mean 6.0)	5.25–6.25 (mean 5.7)	2.03–2.94 (mean 2.52)	0.30–0.43 (mean 0.37)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	very long, with small bulge at proximal end	absent	very long, tapering towards proximal end	absent
L. parahyangensis sp. nov.	14.7–16.9 (mean 15.7)	8.4–9.3 (mean 8.7)	6.5–7 (mean 6.8)	1.69–1.94 (mean 1.80)	0.39–0.45 (mean 0.42)	sharply keeled	brownishcorneous	scaly processes sparse, at periphery	not discernible	?	?	?	?
L. nusakambangensis sp. nov.	11.9–15.0 (mean 13.5)	6.6–9.1 (mean 7.9)	5.25–6 (mean 5.7)	1.65–1.83 (mean 1.72)	0.32–0.40 (mean 0.36)	keeled	brownishcorneous	scaly processes sparse, at periphery	indistinct	with small globular bulge at proximal end	absent	distal part slightly crenated, proximal part long	absent
L. petrukensis sp. nov.	8.1–9.8 (mean 9.0)	5.2–6.3 (mean 5.5)	4.75–5.5 (mean 5.1)	1.49–1.79 (mean 1.62)	0.24–0.37 (mean 0.27)	sharply keeled	brownish- corneous	scaly processes all over teleoconch	indistinct	divided by narrowing into long, cylindrical distal part and slightly dilated proximal part	absent	slightly crenated, abruptly tapering at proximal end	absent
L. tholiformis sp. nov.	13.0–14.4 (mean 13.9)	7.4–7.8 (mean 7.7)	6–6.25 (mean 6.1)	1.77–1.88 (mean 1.82)	0.32–0.40 (mean 0.35)	sharply keeled	brownish- corneous	scaly processes all over teleoconch	indistinct	slightly broader proximally	short	crenated, slightly longer than epiphallus	absent
L. abdidalem sp. nov.	13.1–16.4 (mean 14.0)	6.3–7.8 (mean 6.9)	5.75–6 (mean 5.9)	1.93–2.14 (mean 2.03)	0.33–0.48 (mean 0.38)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	long and narrow, without proximal buldge	absent	crenated, shorter than epiphallus	absent
												long, broadest in
L. madurensis sp. nov.	11.4–12.0 (mean 11.6)	5.5–6.5 (mean 5.9)	5.5–5.75 (mean 5.7)	1.75–2.11 (mean 1.98)	0.38–0.40 (mean 0.39)	sharply keeled	brownish- corneous	scaly processes protruding into hairs	no spiral lines	long, divided into longer, broader distal and shorter, narrower proximal part	absent	middle part, gradually tapering towards proximal end, slightly crenated	absent
L. sewuensis sp. nov.	11.4–16.3 (mean 13.7)	6.3–8.7 (mean 7.2)	5.25–6 (mean 5.7)	1.61–2.25 (mean 1.90)	0.30–0.40 (mean 0.36)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	with globular swelling at proximal end	absent	broad, with two rows of tubercles	absent
L. sukoliloensis sp. nov.	10.4–12.3 (mean 11.2)	6.0–7.4 (mean 6.5)	5–5.75 (mean 5.5)	1.60–1.93 (mean 1.72)	0.22–0.41 (mean 0.29)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	slightly dilated at proximal end	absent	abruptly tapering at proximal end, not distinctly crenated	absent
L. tonywhitteni sp. nov.	14.3–17.6 (mean 16.2)	7.0–8.5 (mean 7.5)	5.75–6 (mean 5.8)	1.93–2.34 (mean 2.15)	0.32–0.41 (mean 0.37)	sharply keeled	brownish- corneous	radial scaly processes	indistinct	divided into long, cylindrical distal part and dilated proximal part	absent	short, broadest at base, not distinctly crenated	present
L. nodifera sp. nov.	14.7–16.1 (mean 15.2)	6.4–8.1 (mean 7.2)	5–5.75 (mean 5.5)	1.93–2.29 (mean 2.12)	0.26–0.38 (mean 0.32)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	divided into cylindrical distal and dilated proximal part	present	moderately long, tuberculated	present
L. pacitanensis sp. nov.	10.6–14.7 (mean 13.1)	6.2–8.2 (mean 7.1)	5.25–6 (mean 5.7)	1.57–2.18 (mean 1.84)	0.28–0.34 (mean 0.31)	sharply keeled	brownish- corneous	scaly processes sparse, at periphery	indistinct	divided by narrowing into long distal and short proximal section	present	moderately long, not crenated	absent
L. intumescens	11.6–14.1 (mean 13.2)	6.1–8.7 (mean 7.5)	5–5.75 (mean 5.3)	1.50–1.98 (mean 1.76)	0.22–0.34 (mean 0.27)	sharply keeled	brownish- corneous	without scaly processes	indistinct	divided by narrowing into long distal and short proximal section	absent	distal part crenated, proximal part short	absent
L. moussoniana	10.2–12.0 (mean 11.1)	5.5–6.7 (mean 6.3)	5–6.25 (mean 5.7)	1.73–1.81 (mean 1.77)	0.37–0.38 (mean 0.38)	keeled	brownish- corneous	?	indistinct	?	?	?	?
L. schepmani	10.5–10.9	6.1–6.2	5–5.25	1.72–1.76	0.21	sharply keeled	brownish- corneous	no scales discernible	distinct	?	?	?	?
L. monticola	8.5–9.5 (mean 9.0)	5.3–6.0 (mean 5.8)	4.5–5.25 (mean 4.8)	1.49–1.60 (mean 1.56)	0.18–0.25 (mean 0.21)	keeled	yellowish or brownish- corneous	no scales discernible	distinct	?	?	?	?
												slightly crenated,
L. smimensis	6.3–11.9 (mean 8.2)	4.1–8.4 (mean 5.6)	4.5–5.5 (mean 4.9)	1.28–1.64 (mean 1.48)	0.18–0.32 (mean 0.25)	angular or rounded	brownish- corneous	hair-like processes	indistinct	long, gradually tapering towards its proximal end	absent	gradually tapering towards proximal end	absent
L. leucochila	16.3–17.5	8.5–9.5	5.25	1.92	0.30	angular	yellowish	scaly processes all over teleoconch	indistinct	?	?	?	?
L. zonifera sp. nov.	12.5–13.2 (mean 12.8)	6.9–7.6 (mean 7.2)	5.5–5.75 (mean 5.6)	1.73–1.81 (mean 1.77)	0.40	keeled	corneous with brownish zone at periphery	scaly processes sparse, at periphery	indistinct	slightly dilated at proximal end	absent	slightly crenated, more than twice as long as epiphallus	absent
L. ciliocincta	12.4–15.6 (mean 13.6)	6.0–8.3 (mean 7.0)	5–6 (mean 5.4)	1.86–2.12 (mean 1.94)	0.22–0.30 (mean 0.27)	keeled	whitish- corneous with brownish band at periphery	scaly processes sparse, at periphery	indistinct	with small bulge at proximal end	absent	short, very broad, tapering towards proximal end	absent
												moderately
							whitish,					long, almost
L. pakidulan sp. nov.	12.5–14.0 (mean 13.4)	6.9–8.1 (mean 7.4)	5.25–5.5 (mean 5.4)	1.70–1.86 (mean 1.81)	0.24–0.30 (mean 0.27)	angular	with reddish- brown band at	without scaly processes	indistinct	short, with large bulge at proximal end	absent	smooth, gradually tapering near	absent
							periphery					proximal
												end
							whitish,
							with
L. dharmai sp. nov.	9.9	7.2	5.5	1.39	0.18	angular	reddish- brown	not observed	indistinct	?	?	?	?
							band at
							periphery
							whitish-
							corneous,
							with one
L. davini	11–14.8	7.1–10.3	5–5.5	1.41–1.67	0.29	angular	reddish- brown band	without scaly processes	indistinct	?	?	?	?
							above and
							one below
							periphery
							whitish,
L. menorehensis sp. nov.	12.4–14.5 (mean 13.4)	8.3–10.4 (mean 9.3)	5.25–5.75 (mean 5.5)	1.31–1.51 (mean 1.44)	0.16–0.24 (mean 0.19)	rounded	with distinct brown band above	without scaly processes	indistinct	short, distal part cylindrical, proximal part slightly dilated	absent	long, broad	absent
							periphery