Redescription and Revised Classification Status of Abbreviata phrynocephali comb. nov. (Spirurida: Physalopteridae) from the Yarkand Toad-Headed Agama Phrynocephalus axillaris (Lacertilia: Agamidae) Author Yan, Xiao-Fei Key Laboratory of Extreme Environmental Biology and Ecological Adaptation & Evolution, College of Life Sciences, Xinjiang Agricultural University, Urumqi 830052, China. Author Wang, Tao Key Laboratory of Extreme Environmental Biology and Ecological Adaptation & Evolution, College of Life Sciences, Xinjiang Agricultural University, Urumqi 830052, China. Author Zhao, Yi-Yang College of Veterinary Medicine, Xinjiang Agricultural University, Urumqi 830052, China. Author Chen, Sheng-Tao Key Laboratory of Extreme Environmental Biology and Ecological Adaptation & Evolution, College of Life Sciences, Xinjiang Agricultural University, Urumqi 830052, China. Author Wang, Yi-Ni Key Laboratory of Extreme Environmental Biology and Ecological Adaptation & Evolution, College of Life Sciences, Xinjiang Agricultural University, Urumqi 830052, China. text Zootaxa 2025 2025-05-12 5633 2 336 350 https://doi.org/10.11646/zootaxa.5633.2.6 journal article 309884 10.11646/zootaxa.5633.2.6 dbd4a274-9159-44c0-90c0-9e543220c072 1175-5326 15435201 DF928AA8-1032-46A1-B86A-EE3E3DBAA577

TABLE 2. Comparison of morphological characters of nematodes of the genus Abbreviata (measurements in mm).

	A. phrynocephalus		A. levicauda		A. tumidocapitis		A. pilbarensis		A. anomala		A. glebopalmae		Pentadentoptera phrynocephali ✱		A. bancrofti		A. borneensis		A. baltazardi		A. adonisi		A. hastaspicula		A. perenticola		A. achari
......continued on the next page BL body length, MW maximum width, ML length of muscular esophagus, GL length of glandular esophagus, NC distance of nerve ring to cephalic end, EPC distance of excretory pore to cephalic end, DC distance of deirids to cephalic end, SL length of left spicule, SR length of right spicule, TL length of tail, ES size of eggs, PP number of pedunculate papillae, PSP number and arrangement of caudal sessile papillae, and UB number of uterine branches. * This species was included for comparison with Abbreviata spp. Please refer to the main text for the proposed taxonomic revisions.
BL body length, MW maximum width, ML length of muscular esophagus, GL length of glandular esophagus, NC distance of nerve ring to cephalic end, EPC distance of excretory pore to cephalic end, DC distance of deirids to cephalic end, SL length of left spicule, SR length of right spicule, TL length of tail, ES size of eggs, PP number of pedunculate papillae, PSP number and arrangement of caudal sessile papillae, and UB number of uterine branches.
Source	This study		Jones (1983)		Jones (1983)		Jones (1986)		Jones (1986)		Jones (1988)		Tao (1985)		Jones, 2013		Schad, 1959		Chabaud, 1953		Sulahian, 1968		Jones, 1979		Mus, 1985		Soota, 1969
Host	Phrynocephalus axillaris		Varanus rosenbergi		Varanus gouldii		Pogona minor mitchelli		Pogona minor minor		Varanus glebopalma		Phrynocephalus axillaris		Phyllurus platurus		Varanus rudicollis		Phrynocephalus helioscopus		Agatna stellio		Aspidites melanocephalus		Varanus giganteus		Calotea versicolor
Sex	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female	male	female
BL	5.58–9.55	6.56–12.57	11.5–18.0	18.5–25.0	8–10.5	11–14	8.9–10.5	13.2–21.0	17.8–20.5	40.4–51.8	8.1–10.7	11.3–13.4	5.5–6.5	6–8	13.4–17.6	23.8	11.00– 24.50	24.00– 37.00	8.50	12.50	16.2	25–32.7	11.17– 12.37	17.62– 30.75	16.0–22.0	26.0–31.0	11.44– 13.31	19.2–21.2
MW	0.37–0.45	0.44–0.61	0.35–0.44	0.50–0.74	0.28–0.38	0.42–0.49	0.30–0.40	0.52–0.67	0.44–0.50	0.56–0.88	0.29–0.31	0.40–0.48	0.274– 0.434	0.308– 0.480	0.428– 0.528	0.64	-	-	0.360	0.475	3.05	0.53–0.62	0.40–0.67	0.66–1.10	0.38–0.62	0.60–0.76	0.33–0.35	0.43–0.45
ML	0.22–0.32	0.18–0.30	0.31–0.37	0.35–0.45	0.36–0.43	0.32–0.53	0.25–0.35	0.28–0.42	0.23–0.29	0.35–0.56	0.39–0.50	0.46–0.51	0.205– 0.226	0.251– 0.297	0.28–0.32	0.32	0.22–0.38	0.30–0.47	0.240	0.210	0.34	0.45–0.54	0.26–0.41	0.37–0.44	0.26–0.40	0.32–0.44	-	-
GL	1.27–1.54	1.46–2.06	2.07–2.50	2.37–3.25	2.10–2.84	2.16–3.04	1.62–1.92	2.20–2.75	2.30–3.10	3.30–4.40	2.40–0.76	0.80–3.50	1.154– 1.497	1.439– 1.731	2.30–2.80	3.26	-	-	1.40	1.80	3.23	3.27–3.84	1.75–2.84	2.22–3.61	2.56–3.26	3.30–4.10	-	-
NC	0.18–0.24	0.16–0.27	0.31–0.37	0.31–0.45	0.27–0.39	0.26–0.39	0.25–0.31	0.26–0.38	0.46–0.59	0.26–0.30	0.32–0.40	0.38–0.44	0.160– 0.205	0.213– 0.240	0.28	0.344	0.24–0.37	0.33–0.42	0.21	0.24	0.27	0.31–0.34	0.23–0.38	0.36–0.43	0.32–0.38	0.33–0.45	-	-
EPC	0.37–0.65	-	0.58–0.68	0.63–0.87	0.49–0.66	0.42–0.67	0.52–0.65	0.48–0.76	0.32–0.55	0.36–0.72	0.78–0.90	0.72–0.84	0.313– 0.326	0.333– 0.366	0.480– 0.568	0.584	0.37–0.60	0.62–0.76	0.30	0.380	-	0.47–0.57	0.50–0.77	0.64–0.80	0.48–0.78	0.44–0.80	-	-
DC	0.37–0.53	0.46–0.55	0.57–0.67	0.63–0.86	0.32–0.51	0.35–0.62	0.45–0.59	0.45–0.76	0.36–0.46	0.38–0.60	0.34–0.48	0.36–0.46	0.166– 0.313	0.320	0.40–0.48	0.624	0.34–0.53	0.54–0.68	0.32	0.370	-	–	0.43–0.61	0.56–0.70	0.39–0.62	0.40–0.66	-	-
VC/BL	-	17.19– 30.22	-	18–31.8	-	22.6–30.0	-	25.9–38.7	-	3.0–4.8	-	21.1–26.9	-	21.32– 23.99	-	21.4	-	27.03– 40.00	-	20.55	-	-	-	21.68- 24.39	-	-	-	20.00– 21.77
SL	0.32–0.53	-	0.50–0.60	-	0.442– 0.594	-	0.325– 0.425	-	0.72–0.90	-	0.48–0.60	-	0.14–0.24	-	1.20–1.36	-	1.94–2.40	-	0.32	-	0.37	-	0.62–0.70	-	0.62–0.80	-	0.28–0.39	-
SR	0.18–0.23	-	0.30–0.34	-	0.197– 0.276	-	0.15–0.21	-	0.22–0.26	-	0.24–0.32	-	1.714– 2.355	-	0.312– 0.320	-	0.29–0.94	-	0.22	-	0.14	-	0.59–0.67	-	0.48–0.56	-	0.08–0.11	-
TL	0.48–0.61	0.29–0.41	-	0.38–0.62	-	0.21–0.30	-	0.30–0.52	-		0.44–0.56	0.19–0.3	-	0.331– 0.365	-	0.520	-	0.19–0.50	0.460	-	-	-	-	0.28–0.43	0.74–0.88	0.24–0.38	0.28–0.42	-
ES (μm)	-	30–38× 46–64	-	32.5–33.7 × 47.5–50	-	31–33× 47–49	-	40–45× 50–60	-	44–48× 72–76	-	28–32× 35–36	-	34–40 × 45–60	-	32 ×50	-	21–29× 36–54	-	30 × 65	-	27–32× 51–62	-	31–39× 31–42	-	32–36 × 36–40	-	22 × 44–55
PP	8	-	8	-	8	-	8	-	10	-	8	-	8	-	8	-	8	-	10	-	6	-	14	-	8	-	8	-
PSP	13	-	7	-	7	-	13	-	11	-	7	-	13	-	7	-	11	-	11	-	8	-	7	-	7	-	12	-
UB	-	6	-	4	-	4	-	1	-	4	-	4	-	6	-	4	-	4	-	4	-	4	-	4	-	4	-	4