A revision of the Metaphire peguana species-group (Oligochaeta: Megascolecidae) from Vietnam Author Nguyen, Tung T. Department of Biology, School of Education, Can Tho University, Vietnam. Author Lam, Dang H. 0000-0002-1236-1087 Department of Biology, School of Education, Can Tho University, Vietnam. & lamhaidangct @ gmail. com; https: // orcid. org / 0000 - 0002 - 1236 - 1087 lamhaidangct@gmail.com Author Nguyen, Anh D. 0000-0001-9273-0040 Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay, Hanoi. & ducanh 410 @ yahoo. com; https: // orcid. org / 0000 - 0001 - 9273 - 0040 & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay, Hanoi. ducanh410@yahoo.com text Zootaxa 2023 2023-03-15 5255 1 113 135 http://dx.doi.org/10.11646/zootaxa.5255.1.15 journal article 54940 10.11646/zootaxa.5255.1.15 694336af-f170-49b8-8f98-2788d5a759d8 1175-5326 7744555 3F27E81A-87C7-48CD-8DF7-6FFDA0CD4907

TABLE 3: A morphological comparison between Metaphire peguana -group members and M. pacseana

No.	Characters	M. bahli	M. doiphamon	M. dorsomultitheca	M. kiengiangensis	M. nhuongi	M. peguana peguana	M. peguana laisonensis	M. pacseana
1.	Length (mm)	76–170	101–236	63–107	115–286	206–266	81–157	53–114	273–328
2.	Diameter average (mm)	3,02–5.43	4.92–8.44	2.97–4.53	4.57–8.36	8.59–9.08	3.39–6.27	2.44 –4.46	6,14–11,41
3.	Segments	57–135	78–115	75–122	66–152	118–145	60–157	62–118	128–163
4.	First dorsal pore	12/13	12/13	12/13	12/13	12/13	12/13 or 13/14	12/13	12/13
5.	Setae between male pores	0–10	3–16	3–6	10–19	12–15	2–13	4–13	17–50
6.	Spermathecal pore	3 pairs, 6/7/8/9	3 pairs, 6/7/8/9	Numerous, 7/8/9	Numerous, 6/7/8/9	4 pairs, 5/6/7/8/9	3 pairs, 6/7/8/9	3 pairs, 6/7/8/9	3 pairs, 6/7/8/9
7.	GM in spermathecal region	Absent	4 pairs in vi–ix	Absent	Absent	Absent	Absent	Absent	Absent
8.	GM in male region	2 pairs, 17/18/19	2 pairs, 17/18/19	Absent	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, xvii, xix	2 pairs, 17/18/19
9.	Male GM position	Inline with male pores	Inline with male pores	Invisible from outside	Inline with male pores	Inline with male pores	Inline with male pores	Inline with male pores	Near mid–ventral line
10.	Male GM shape	Invaginated, slit like openings	Not invaginated, disc shaped	Invisible from outside	Invaginated	Not invaginated, disc shaped	Not invaginated, disc shaped	Invaginated, slit like or ellipse- shaped openings	Not invaginated, elipsoidal
11.	Distance between male pores/body circumference	0.11–0.23	0.21–0.31	0.18–0.25	0.20–0.35	0.27–0.32	0.22–0.38	0.21–0.33	0.22–0.40
12.	State of septum 10/11	Only appear ventrally	Only appear ventrally	Only appear ventrally	Only appear ventrally	Only appear ventrally	Only appear ventrally	Only appear ventrally	Fully developed
13.	Spermatheca	Bithecate	Bithecate	Polythecate	Polythecate	Bithecate	Bithecate	Bithecate	Bithecate
14.	Caeca	Simple	Simple or serrate	Simple	Simple	Simple	Simple	Simple	Manicate
15.	Testis sacs	Separated	Separated or connected	Separated	Separated	Separated	Separated	Separated	Connected
16.	Prostatic gland	xvi – xxii	xvi–xx	xvii–xx	xvii–xx	xvii–xix	xvi–xxi	xvii–xx	xvi–xxiii
17.	Base of diverticulum	Ampulla base	Ampulla base or on ampulla duct	Ampulla base	Ampulla base	Ampulla base	Ampulla base	On ampulla duct	Ampulla base
18.	Accessory gland	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, 17/18/19	2 pairs, xvii, xix	2 pairs, 17/18/19
19.	Accessory gland structure	Muscular–walled cover	Muscular–walled cover	Muscular–walled cover	Muscular–walled cover	Muscular–walled cover	Muscular–walled cover	Muscular–walled cover	Massed