Three new species of earthworms belonging to the genera Amynthas and Metaphire (Oligochaeta: Megascolecidae) from Yunnanı China Author Yuan, Zhu Author Dong, Yan Author Jiang, Jibao Author Qiu, Jiangping text Journal of Natural History 2019 2019-10-28 53 31 1961 1974 journal article 24130 10.1080/00222933.2019.1680760 959b13ea-8bec-4b77-b018-e50130773cf3 1464-5262 3654511

Table 4. A comparison of characters of Metaphire daliensis sp. nov., Metaphire californica (Kinberg, 1867), Metaphire leonoris (Chen, 1946) and Metaphire insulana (Gates, 1930).

Character	M. daliensis sp. nov.	M. californica	M. leonoris	M. insulana
Body length * width (mm)	110-143 * 3.9–7.2	50-156 * 3.0–5.0	100-170*5.0–7.0	79*5.0
Pigment
Dorsum	No pigment before VIII, others present	Dark red-brown	Colour greyish purple, chocolate on	Bluish-grey anterior to the clitellum,
	ochre		anterior dorsum	reddish-brown posterior to the
				clitellum
Ventrum	No pigment anterior to segment VIII, ochre	?	Pale	Greyish
in posterior segments
First dorsal pore	11/12	11/12	12/13	12/13
Setal number in segment
Between male pores	22	14-20	11	16
Between spermathecal	26(VIII)	38-41(VII)	25(IX)	32(VIII)
pores
Spermathecal pores	Two pairs, in 7/8-8/9, about 1/3	Two pairs, in 7/8-8/9,	Two pairs, in 7/8-8/9, about 1/3-1/2	Two pairs,in 7/8-8/9,slightly more than
	circumference apart ventrally,	about 1/3-1/2	circumference apart ventrally, each in an	one-half of the circumference apart
	the second pair turns outward obvious	circumference apart	ellipsoid glandular depression, genital
		ventrally, no genital	papilla lacking,sometimes present on VII
		papillae in the	and VIII ventral
		preclitellar region
Male pores	About 1/3 circumference ventrally apart	About 2/5 circumference	In deep pouches, about 1/2 circumference	On XVIII internal to the lines of the
	from each other,each located in the oval	ventrally apart from	ventrally apart from each other, genital	spermathecal pores,at the bottom of
	copulatory pouches, and a slightly	each other, in lateral	papillae 1–5 usually found in each other	small depressions in the setal circle,
	extended mastoid on the top,	slits of retracted		about in line with setae, no
	surrounded with 3–5 circular folds. In	copulatory pouches		copulatory papillae
the copulatory chamber and male pores
have 1–2 papillae (two left, one right)
Intestinal caeca	From XXVIII to XXVII, between simple and	From XXVI or XXVII to	From XXVII to XXII, simple, smooth on both	From XXVII to XXIV, the dorsal margins
	complex	XXIII or XXV, simple	edges	slightly incised
Spermathecae	Ampulla heart-shaped, about 3.0 mm,	Ampulla large, heart-	Ampulla either heart-shaped or sac-like,	Ampulla duct is shorter than the
	ampulla duct short and thick, the length	shaped, ampulla duct	about 3.5–5.0 mm long, ampulla duct	irregularly ovoid ampulla, the
	of diverticulum about 3.0 mm, terminal	stalk short and stout,	shorter, diverticulum shorter than main	diverticulum is longer than twice the
	1/2 slightly dilated into an elongated	diverticulum tubular,	pouch, its terminal 2/3 twisted in zigzag	combined length of duct and
	oval-shaped seminal chamber	slender, coiled	fashion, with a round terminal knob	ampulla and is twisted into a coiled
				mass
Prostate glands	In XVII-XIX, underdeveloped	In XVIII, racemose	In XVI-XXI or XXII, developed	In XV-XX on the right side, XVI-XXI on
				the left side
Accessory glands near	Invisible	Invisible	In large masses, sessile	Invisible
prostate