Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae) Author Lowe, Graeme Author Kovařík, František text Euscorpius 2019 2019-12-31 281 1 128 https://doi.org/10.18590/euscorpius.2019.vol2019.iss281.1 journal article 157784 10.18590/euscorpius.2019.vol2019.iss281.1 67120029-de04-4f6e-b3e8-460ef3896e70 1536-9307 7143610 FEBA0106-02A3-4465-8D39-9FF32634EEF

Table 1. Character states for species examined. States of 5 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of trichobothrium d 2 on pedipalp femur; ratio R 123 = d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀); and size and shape of female basal pectinal tooth.

SPECIES Grosphus goudoti Grosphus hirtus Grosphus madagascariensis Grosphus voahangyae Neogrosphus griveaudi	femur d 2 carinal internal/carinal internal/carinal carinal internal/ dorsal?	R 123 0.82 0.32–0.83 0.45–1.04 0.44–0.64 0.55	PTC ♂ 19–22 17–20 15–20 15–19 29–31	PTC ♀ 17–19 14–18 13–18 14–16 27–29	basal pectine tooth ♀ short, wide short, wide short, wide short, wide short, wide	hemisperm. basal lobe distal distal proximal distal distal	hemisperm. posterior lobe long long long long short	spiracles elliptic ovoid elliptic ovoid narrow elliptic	met. I ventro- med. carinae granulate granulate granulate granulate granulate	telotarsus setation sparse sparse sparse sparse dense	UV fluorescence weak weak weak weak strong
Teruelius ankarafantsika	dorsal	<0.50	27–31	24–27	long, narrow	distal	short	narrow	smooth	dense	strong
Teruelius ankarana	dorsal/carinal	<0.50	36–41	31–35	long, narrow	distal	short	narrow	smooth	dense	strong
Teruelius annulatus	dorsal	<0.50	32–34	24–29	long, narrow	distal	short	narrow	smooth	dense	strong
Teruelius bistriatus	dorsal	<0.50	27–30	24–25	long, narrow	–	–	narrow	smooth	dense	strong
Teruelius feti	dorsal	<0.50	30	25–28	long, narrow	–	–	narrow	smooth	dense	strong
Teruelius flavopiceus	dorsal	<0.50	27–32	24–30	long, narrow	distal	short	narrow	crenulate/ smooth	dense	strong
Teruelius grandidieri	dorsal	<0.50	34–40	30–34	long, narrow	distal	short	narrow	crenulate/ smooth	dense	strong
Teruelius intertidalis	dorsal	<0.50	32–34	28–30	long, narrow	distal	short	narrow	smooth	dense	strong
Teruelius limbatus	dorsal/carinal	<0.50	25–32	23–29	long, narrow	distal	short	narrow	crenulate/ smooth	dense	strong
Teruelius mahafaliensis	dorsal	<0.50	28–40	27–31	long, narrow	distal	short	narrow	crenulate/ smooth	dense	strong
Teruelius olgae	dorsal	<0.50	29–33	26–29	long, narrow	short	short	narrow	weakly crenuate	dense	strong