Antarctic Kinorhyncha: Seven new species from the Antarctic Peninsula Author Sørensen, Martin V. 4143D650-12FC-4914-93F5-2C39339A7156 Natural History Museum of Denmark, University of Copenhagen, 2100 Copenhagen, Denmark. mvsorensen@snm.ku.dk Author Macheriotou, Lara 1037C143-8562-41A3-8149-BC3DE4323845 Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. lara.macheriotou@ugent.be Author Braeckman, Ulrike 6C48F169-ECC8-4F98-A9CF-9B2A664DA903 Marine Biology Research Group, Department of Biology, Ghent University, 9000 Ghent, Belgium. Ulrike.Braeckman@ugent.be Author Smith, Craig R. F004E0AC-049A-48D6-9369-EBE18ECEF53C Department of Oceanography, University of Hawai’i at Mānoa, Honolulu HI 96822, Hawaii, USA. craigsmi@hawaii.edu Author Ingels, Jeroen F6107F36-C67F-4CB8-A6C0-DFBF66D3C464 Coastal and Marine Laboratory, Florida State University, St. Teresa FL 32346, Florida, USA. & National Institute for Water and Atmospheric Research, Wellington 6021, New Zealand. jeroen.ingels@niwa.co.nz text European Journal of Taxonomy 2025 2025-07-01 1000 1 1 102 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2947/13307 journal article 310863 10.5852/ejt.2025.1000.2947 7b56d2d4-1394-4a8a-9448-547bd293786d 2118-9773 15808384 98563124-EFCC-4542-B5AB-E14C0C3978DD

Table 24. Species with the three most common echinoderid dorsal patterns of glandular cell outlets type 1 on segments 1 to 9, listed together with the middorsal spine pattern (MDS) for each species. The table is updated from Sørensen et al. (2020), and names in boldface mark newly added species. In each column species of already recognised or newly proposed species groups are listed together, followed alphabetically by additional species that so far have not been assigned to a species group.

	MD seg. 1–3 PD seg. 4–9		MD seg. 1–3 PD seg. 4–9		MD seg. 1–3, 5, 7 PD seg. 4, 6, 8–9
1–3 Indicates the segment(s) from which information is missing, although other outlets fit the pattern. ** The species has two middorsal outlets on segment 1.
MDS		MDS		MDS
4-8	Cephalorhyncha nybakkeni (Higgins, 1986)		E. coulli species group:		E. horni species group:
		4	E. cyaneafictus Cepeda et al., 2022	–	E. horni Higgins, 1983
	E. coulli species group:	–	E. inaequalis Herranz et al., 2023	–	E. parahorni Cepeda et al. 2019
4	E. blazeji Grzelak & Sørensen, 2022 1	4	E. marthae Sørensen, 2014	–	E. wilberti Anguas-Escalante et al., 2023
4	E. angelae Cepeda et al., 2022 2	4	E. ohtsukai Yamasaki & Kajihara, 2012
		4	E. regina Yamasaki, 2016	4,6,8	Echinoderes abbreviatus Higgins, 1983
	E. dujardinii species group:	4	E. serratulus Yamasaki, 2016	4,6,8	E. ahlfeldae sp. nov.
4-8	E. chandrasekharai Sørensen et al., 2020			4,6,8	E. anniae Sørensen et al., 2018 3
4-8	E. dujardinii Claparède, 1863		E. remanei species group:	4	E. antalyaensis Yamasaki & Durucan, 2018
4-8	E. gerardi Higgins, 1978	4-8	E. angustus Higgins & Kristensen, 1988	4,6,8	E. antarcticus sp. nov.
4-8	E. imperforatus Higgins, 1983	4-8	E. beringiensis Adrianov & Maiorova, 2022	4,6,8	E. apex Yamasaki et al., 2018
4-8	E. kozloffi Higgins, 1977	4-8	E. galadrielae Grzelak & Sørensen, 2022	4,6,8	E. arlis Higgins, 1966
4-8	E. pacificus Schmidt, 1974	4-8	E. quasae Herranz et al., 2024	4,6	E. astridae Sørensen, 2014
4-8	E. sensibilis Adrianov et al., 2002	4-8	E. pennaki Higgins, 1960	4-8	E. aureus Adrianov et al., 2002
4-8	E. songae Sørensen et al., 2020	4-8	E. remanei (Blake, 1930)	4,6,8	E. bermudensis Higgins, 1982
4-8	E. sublicarum Higgins, 1977	4-8	E. romanoi Landers & Sørensen, 2016	4,6	E. bispinosus Higgins, 1982
4-8	E. worthingi Southern, 1914	4-8	E. xalkutaat Cepeda et al., 2019	4,6,8	E. brevipes Cepeda et al., 2019
4-8	Echinoderes sp. Andaman Isl.			4	E. capitatus Zelinka, 1928
		4-8	Cephalorhyncha polunga Sánchez et al., 2019	4,6,8	E. crux sp. nov. 2
		4-8	Echinoderes abeli Anguas-Escalante et al., 2023	4,6	E. dalzottoi Grzelak & Sørensen, 2022
		4-8	Echinoderes adrianovi Herranz et al., 2014	4,6,8	E. goku Rucci et al., 2022
		4-8	E. aquilonius Higgins & Kristensen, 1988	4,6,8	E. hakaiensis Herranz et al., 2018
		4-8	E. augustae Sørensen & Landers, 2015	4,6,8	E. hamiltonorum Sørensen et al., 2018
		4-8	E. australis Sánchez et al., 2024 **	4,6,8	E. higginsi Huys & Coomans, 1989
		4-8	E. barbadensis Cepeda et al., 2019	4,6,8	E. intermedius Sørensen, 2006
		4-8	E. bookhouti Higgins, 1964	4,6,8	E. kathleenhannae sp. nov.
		4-8	E. dubiosus Sørensen et al., 2018 2,3,**	4,6,8	E. leduci Grzelak & Sørensen, 2022
		4-8	E. ferrugineus Zelinka, 1928	4,6,8	E. legolasi Grzelak & Sørensen, 2022
		4-8	E. gama Yamasaki et al., 2020	4,6,8	E. multiporus Yamasaki et al. 2018
		4-8	E. juliae Sørensen et al., 2018	4,6,8	E. nataliae sp. nov.
		4-8	E. kaempfae Yamasaki et al., 2019	4-8	E. okiensis Yamasaki et al. 2024
		4,6,8	E. kajiharai Yamasaki et al., 2020	4,6,8	E. peterseni Higgins & Kristensen, 1988
		4-8	E. kanni Thormar & Sørensen, 2010	4,6,8	E. riceae Herranz et al., 2014
		4-8	E. levanderi Karling, 1954	4-8	E. sanctorum Sanchez et al., 2022
		4-8	E. mamaqucha Grzelak et al., 2021 **	4,6,8	E. shahmaranae Sørensen et al., 2021
		4-8	E. muricatus Pardos et al., 2016	4,6,8	E. skipperae Sørensen & Landers, 2014
		4,6,8	E. riedli Higgins, 1966	4	E. unispinosus Yamasaki et al., 2018
		4,6,8	E. samwisei Grzelak & Sørensen, 2022	4,6	E. uozumii Yamasaki et al., 2020
		4,6,8	E. schwieringae Yamasaki et al., 2019	4,6,8	E. wallaceae Higgins, 1983
		4-8	E. tchefouensis Lou, 1934	4,6,8	E. xiphophorus Adrianov & Maiorova, 2021
		4-8	E. zeppilliae Sanchez et al. 2022	4-6,8	Fissuroderes sorenseni Herranz & Pardos, 2013
		4-8	Fissuroderes higginsi Neuhaus & Blasche, 2006	4,6,8	Meristoderes boylei Herranz & Pardos, 2013
		4-8	F. novaezealandia Neuhaus & Blasche, 2006	4	M. galatheae Herranz et al., 2012
		4-8	Meristoderes taro Sánchez et al., 2019	4,6,8	M. herranzae Sørensen et al., 2013
		4-8	Polacanthoderes grzelakae sp. nov.	4,6,8	M. macracanthus Herranz et al., 2012
		4-8	P. martinezi Sørensen, 2008
		4-8	P. shiraseae Yamasaki et al. 2022